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rowth and form of marine organisms inhabiting hard substrata, the
G"marine sessile organisms," ischaracterized by anumber
ofremarkable properties. One remarkable feature of these organisms
is that many ofthem can be characterizedasmodularorganisms.
Modularorganisms are typically built ofrepeated units, the modules,
which might be a polyp in a coral colony or afrond in seaweeds. In
most cases, the modulehas adistinctive form, while the growth form
of the entire colony is frequently an indeterminate form.
Indeterminategrowthindicatesthatthe same growthprocess mayresult in
an infinite numberofdifferentrealizations ofthe growthform.This
isincontrast to unitaryorganisms such asvertebrates and insects, in
which a single-celled stage develops into a well-defined,
determinate structure. In many cases the growth process in modular
organisms leads to complex shapes, which are often quite difficult
to describe in words. In most of the biological literature these
forms are only described in qualitativeand rather vague terms, such
as "thinlybranching,""tree-shaped" and "irregularlybranching."
Anothermajor characteristic ofmarine sessile organisms is that
there is frequentlyastrongimpactofthe physical environmenton the
growthprocess, leading to a variety of growth forms. Growth by
accumulation of modules allows the organism to fit its shape to its
environment i.e., have plasticity. In many seaweeds, sponges, and
corals, differences in exposure to water movement cause significant
changes in morphology. Agood example of this plasticity is the
Indo-Pacific stony coral Pocillopora damicornis(Veron and Pichon
1976) shown in Plg.r.i. In very sheltered environments, this
species has a thin-branching growth form. The growth form gradually
transforms to a more compact shape when the exposure to water
movement increases.
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