Carefully examines the events recorded at the major Permo-Triassic boundary sections and documents the patterns of extinction and survival among the major groups of marine and terrestrial plants and animals. Erwin also provides a detailed summary of the climatic, geologic, geophysical and geochemical events of the Late Permian and Early Triassic.

Carefully examines the events recorded at the major Permo-Triassic boundary sections and documents the patterns of extinction and survival among the major groups of marine and terrestrial plants and animals. Erwin also provides a detailed summary of the climatic, geologic, geophysical and geochemical events of the Late Permian and Early Triassic.

Introduction by Stephen Jay Gould

In the geological record, there are five major mass extinctions—the "Big Five." The most famous happened at the end of the Cretaceous Period, when the dinosaurs and two-thirds of all marine animal species were wiped out, opening the door for the age of mammals and the rise of Homo Sapiens. Using this example as a springboard, David M. Raup leaps into an egaging discussion of the theories, assumptions, and difficulties associated with the science of species extinction. Woven is along the way are stories of the trilobite eye, tropical reefs, flying reptiles, and the fate of the heath hen on Martha's Vineyard, a very modern extinction.

This is the first major book to present a comprehensive overview of the current state of extinction studies. At the end of the journey, Raup has put forward the best science of the day to answer the question posed by the title: Bad genes or bad luck?

"An eminently entertaining and informative read." —Malcolm W. Browne, New York Times Book Review

"A delightful little book about life on this planet and about extinctions, in particular. It is as much about the philosophy and methodology of science as about the downside of evolution." —Clark R. Chapman, Planetary Science Institute

"David Raup's Extinction will change the way many of us perceive our world. In a style that is both elegant and persuasive, Raup undercuts the popular and comfortable notions that extinction is a mark of failure. . . . We are shown a world that is less certain, but in many ways more interesting than the one we imagined we occupied." —Roger Lewin, author of Bones of Contention

How can the tracks of dinosaurs best be interpreted and used to reconstruct them? In many Mesozoic sedimentary rock formations, fossilized footprints of bipedal, three-toed (tridactyl) dinosaurs are preserved in huge numbers, often with few or no skeletons. Such tracks sometimes provide the only clues to the former presence of dinosaurs, but their interpretation can be challenging: How different in size and shape can footprints be and yet have been made by the same kind of dinosaur? How similar can they be and yet have been made by different kinds of dinosaurs? To what extent can tridactyl dinosaur footprints serve as proxies for the biodiversity of their makers? Profusely illustrated and meticulously researched, Noah's Ravens quantitatively explores a variety of approaches to interpreting the tracks, carefully examining within-species and across-species variability in foot and footprint shape in nonavian dinosaurs and their close living relatives. The results help decipher one of the world's most important assemblages of fossil dinosaur tracks, found in sedimentary rocks deposited in ancient rift valleys of eastern North America. Those often beautifully preserved tracks were among the first studied by paleontologists, and they were initially interpreted as having been made by big birds-one of which was jokingly identified as Noah's legendary raven.

Illustrated Key to Skulls of Genera of North American Land Mammals is a manual that contains illustrations of North American land mammals such as marsupials, shrews, bats, moles among many others. This manual is a well-illustrated key, useful for identifying mammals through cranial characteristics. It also contains line-drawings, and many photographs to aid in identifying related genera. The distribution, diversity, and characteristics of each order and family of land mammals found in North American and to the north of Mexico are briefly discussed. J. Knox Jones, Jr., has been a practicing mammalogist for more than 40 years. Currently he is a Paul Whitfield Horn Professor of Biological Sciences at Texas Tech and a Curator in the Museum there. Jones has authored or edited 14 books among is more than 350 publications, and has studied mammals on five continents. He is a past president of the American Society of Mammalogists and has been awarded the C. Hart Merriam Award, the H. H. T. Jackson Award, and Honorary Membership by that society. In 1992, he was selected as Texas Distinguished Scientist of the Year by the Texas Academy of Science, and was awarded the Donald W. Tinkle Research Excellence Award by the Southwestern Association of Naturalists. Richard W. Manning is a member of the faculty of Southwest Texas State University in San Marcos. He has authored more than 40 publications, most of which deal with mammals. Manning has had considerable instructional experience in laboratories in mammalogy, and has been cited for his excellence in teaching. He is also an avid field biologist, and thus has studied mammals in their natural habitats as well. Manning took most of the photographs used in this laboratory manual and made many of the line drawings.

This edited volume explores the various views on the origins of tetrapods-amphibians, reptiles, birds, and mammals-views that agree or differ depending in part on how certain fossil animals are classified and which methodology is used for classification. Eighteen chapters by an international group of paleontologists and neontologists here present current hypotheses, emphasizing the kinds of data needed to answer controversial questions, as well as the variety of solutions that emerge from diferent analyses of the same data set. The book is arranged in five sections, each of which contains an overview essay that either describes the development of various schools of thought regarding the origin of the tetrapod group in question or critically summarizes the arguments presented in the section. The first section addresses the origins of tetrapods as a group, focusing on lobe-finned fishes and early tetrapods. Next is a section dealing with amphbians, followed by one on reptiles. The fourth section concerns avian origins, and the final section treats the origins and early diversification of mammals. With an overall goal of stimulating critical evaluation by the reader rather than providing unequivocal answers, this volume will be of particaular interest to vertebrate paleontologists, evolutionary morphologists, and ichthyological, herpatological, avian, and mammalian systematists.

Essays discuss size, time, the landscape, the world view of modern biology, two-dimensional animals, sabertooth tigers, evolution, and the scientific method.

The authors argue that the growth pattern and form of the colony in many bryozoans is an adaptive strategy rather than a stable genetic character. "Bryozoan Evolution is profusely illustrated and has a bibliography of over 400 titles. It will find an appreciative audience of paleontologists, invertebrate zoologists, and ecologists thanks to its innovative and detailed evaluations of the roles of ecology, adaptive and functional morphology, life histories, biomechanics, developmental constraints, and chance on the evolution of the marine taxa of this speciose group."--Russel L. Zimmer, Science "This book is an excellent source of information on the functional morphology and variety of colonial architecture in bryozoans, very well illustrated, and worth reading at least twice."-Robert L. Anstey, Paleobiology "Even as one of the converted, I found the book a stimulating combination of paleobiology and ecology. In many ways it is a 'teaser'-the authors suggest a number of interesting hypotheses, and can test only some of them. Perhaps most important, McKinney and Jackson provide a plethora of fascinating ideas and examples that demonstrate the potential of this group of animals, and that should stimulate more work."-Michael S. Keough, TREE "This stimulating book is sure to promote further interest in bryozoans. It will appeal to biologists and paleontologists alike."-Paul Taylor, Times Higher Education Supplement

In this volume the author examines the pervasive pattern of alternation of structure and height of vertebral neural spines in the context of a larger review of axial structure in the most terrestrial of primitive Permo-Carboniferous tetrapods. He concludes that a coupled pattern of axial dorsiflexion and rotation played a significant role in primitive terrestrial locomotion.

Assembles Kurten's seminal papers in one volume, not only making them available once again but at the same time presenting a number of concepts and methodological innovations as they were first conceived.

How does bison meat taste after being frozen for 30,000 years? Were Ice Age cave painters trying to create "art" or just record history? How did ancient oil spills occur, before there were oil companies to create them? Those are just some of the questions renowned paleontologist Bjorn Kurten answers in this collection of lighthearted essays on fossils, ancient life, and related topics. Written for the general reader, these lively pieces range from a look at how scientific theories are created to some new views of old myths. Among the topics Kurten examines are the history of the Mediterranean Sea, the origin of birds, the theory of plate tectonics (continental drift), and the discovery of Piltdown Man, the "missing link" fossil forgery that fooled scientists for more than 40 years.

And, true to its title, the book offers a humorous "recipe" for freezing a mammoth that is tundra-tested, if not totally foolproof. "You may have to expend a few hundred mammoths before everything works out," the reader is cautioned, "But there are plenty of them." (Although the author hasn't tasted the fruits of his mammoth recipe, he did feast on some ancient bison meat that dated from 30,000 years ago. Kurten described the taste as "agreeable.")

Throughout these essays Kurten brings the prehistoric world alive with enthusiasm and humor, emphasizing that paleontology is the study of those that lived long ago instead of those who are long dead. As he says, "Isn't it more fun to see a dinosaur as something that used to live, rather than as the monstrous heap of bones which it happens to be at present?"

John H. Ostrom's expeditions to the Bighorn Basin of Wyoming and Montana in the 1960s resulted in discoveries and research that would change long-held concepts in paleontology. This fiftieth-anniversary edition of his now well-known description of the type specimen of Deinonychus antirrhopus revisits the work that redefined theropod dinosaurs as the intelligent, agile, and gregarious ancestors of modern birds and led in the late twentieth century to a renaissance in the study of dinosaurs and the evolution of flight. Distributed for the Yale Peabody Museum of Natural History

Fish, or lower vertebrates, occupy the basal nodes of the vertebrate phylogeny, and are therefore crucial in interpreting almost every feature of more advanced vertebrates, including amphibians, reptiles, birds and mammals. Recent research focuses on combining evolutionary observations - primarily from the fish fossil record - with developmental data from living fishes, in order to better interpret evolutionary history and vertebrate phylogeny. This book highlights the importance of this research in the interpretation of vertebrate evolution, bringing together world-class palaeontologists and biologists to summarise the most interesting, current and cutting-edge topics in fish evolution and development. It will be an invaluable tool for researchers in early vertebrate palaeontology and evolution, and those particularly interested in the interface between evolution and development.

Approximately 99% of all life that has ever existed is extinct. Fortunately, these long dead species have left traces of their lives and interactions with other species in the rock record that paleoecologists use to understand how species and ecosystems have changed over time. This record of past life allows us to study the dynamic nature of the Earth and gives context to current and future ecological challenges. This book brings together forty-four classic papers published between 1924 and 1999 that trace the origins and development of paleoecology. The articles cross taxonomic groups, habitat types, geographic areas, and time and have made substantial contributions to our knowledge of the evolution of life. Encompassing the full breadth of paleoecology, the book is divided into six parts: community and ecosystem dynamics, community reconstruction, diversity dynamics, paleoenvironmental reconstruction, species interaction, and taphonomy. Each paper is also introduced by a contemporary expert who gives context and explains its importance to ongoing paleoecological research. A comprehensive introduction to the field, Foundations of Paleoecology will be an essential reference for new students and established paleoecologists alike.

During the past forty-three years our Division of Zoology of the Royal Ontario Museum has been most active in collecting various species of animals throughout the Province of Ontario, with particular emphasis on the local fauna of the Toronto region. During this time a number of outstanding naturalists have not only made careful collections of animals, but have also recorded accurate observations on the species with which they were familiar. This publication, we most sincerely hope, will be the first of many similar contributions to the study of the distribution of the fauna of Ontario.

Humans have "gone underground" for survival for thousands of years, from underground cities in Turkey to Cold War-era bunkers. But our burrowing roots go back to the very beginnings of animal life on earth. Without burrowing, the planet would be very different today. Many animal lineages alive now-including our own-only survived a cataclysmic meteorite strike 65 million years ago because they went underground. On a grander scale, the chemistry of the planet itself had already been transformed many millions of years earlier by the first animal burrows, which altered whole ecosystems. Every day we walk on an earth filled with an under-ground wilderness teeming with life. Most of this life stays hidden, yet these animals and their subterranean homes are ubiquitous, ranging from the deep sea to mountains, from the equator to the poles. Burrows are a refuge from predators, a safe home for raising young, or a tool to ambush prey. Burrows also protect animals against all types of natural disasters: fires, droughts, storms, meteorites, global warmings-and coolings. In a book filled with spectacularly diverse fauna, acclaimed paleontologist and ichnologist Anthony Martin reveals this fascinating, hidden world that will continue to influence and transform life on this planet.

Sauropod dinosaurs were the largest animals ever to walk the earth, and they represent a substantial portion of vertebrate biomass and biodiversity during the Mesozoic Era. The story of sauropod evolution is told in an extensive fossil record of skeletons and footprints that span the globe and 150 million years of earth history. This generously illustrated volume is the first comprehensive scientific summary of sauropod evolution and paleobiology. The contributors explore sauropod anatomy, detail its variations, and question the myth that life at large size led to evolutionary stagnation and eventual replacement by more "advanced" herbivorous dinosaurs. Chapters address topics such as the evolutionary history and diversity of sauropods; methods for creating three-dimensional reconstructions of their skeletons; questions of sauropod herbivory, tracks, gigantism, locomotion, reproduction, growth rates, and more. This book, together with the recent surge in sauropod discoveries around the world and taxonomic revisions of fragmentary genera, will shed new light on "nature's greatest extravagances."

The Grand Staircase-Escalante National Monument is the location of one of the best-known terrestrial records for the late Cretaceous. Prior fieldwork confirmed the richness of the area, but a major effort begun in the new century has documented over 2,000 new vertebrate fossil sites, provided new radiometric dates, and identified five new genera of ceratopsids, two new species of hadrosaur, a probable new genus of hypsilophodontid, new pachycephalosaurs and ankylosaurs, several kinds of theropods (including a new genus of oviraptor and a new tyrannosaur), plus the most complete specimen of a Late Cretaceous therizinosaur ever collected from North America, and much more. The research documented in this book is rewriting our understanding of Late Cretaceous paleobiogeography and dinosaur phyletics. At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah is a major stepping stone toward a total synthesis of the ecology and evolution of the Late Cretaceous ecosystems of western North America.

William Diller Matthew (1871-1930), who could be called the Father of Mammalian Paleontology, occupies a major position in the history of North American paleontology. In his biography of this paleontologist's paleontologist, Edwin Colbert paints a compelling portrait of the scientist's life and work. Using personal letters, archives, and accounts from those who knew Will Matthew, Colbert presents a delightful and informative account of Matthew's experiences around the turn of the century, complete with photographs of his excavations, world travels, time in New York, family, and environs. Matthew's unique contribution to science rested on his dual perspective as both geologist and paleontologist. He diligently collected and catalogued fossils while at the same time noting stratigraphic details that gave significant time-related information on these fossils. Matthew wrote widely on paleontology at the turn of the century, but most well known is his extensive essay "Climate and Evolution", written in 1915, which attracted almost immediate attention from the paleontological-zoological community. Characteristic of his no-nonsense approach to his work, this essay remains vital and inspirational to paleontological researchers today. Will Matthew was also keen on reaching the public at large and published profusely in the American Museum Journal (which after 1918 became Natural History magazine), where he described his fossil finds, new exhibits, and other interesting research. He also published several American Museum guide leaflets on specific subjects, such as the horse. His informative description "'The Horse', wrote a small boy in his essay, 'is the square animal with a leg at eachcorner, '" - a definition that obviously charmed Matthew. Peppered with such anecdotes, the engaging and highly readable William Diller Matthew, Paleontologist: The Splendid Drama Observed vividly depicts a man respected and revered by many generations of scientists.

Questo secondo volume prosegue la breve trattazione sulle catastrofi naturali. E la volta delle catastrofi dell acqua e dell aria, spesso sui giornali quando un uragano o un inondazione colpiscono il nostro pianeta. E anche delle minacce cosmiche, rare ma in grado di annientare la nostra civilta.

Vedremo quegli straordinari eventi di cui conosciamo l esistenza dallo studio degli strati geologici e dei fossili, ma la cui causa e ancora avvolta nel mistero. Si tratta delle estinzioni di massa, episodi in cui la biosfera e cambiata in maniera improvvisa e drammatica a causa di una catastrofe globale.

Tra un aneddoto e una digressione sulle basi scientifiche dei fenomeni e passando attraverso qualche caso di studio, si incuriosisce il lettore su un tema importante e coinvolgente.

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Die Oberflache der Erde hat eine bewegte Geschichte hinter sich. So stabil uns manche grossraumigen Landschaftsstrukturen wie Gebirgszuge, Tiefebenen und Ozeane auch erscheinen mogen, so sehr hat sich das Antlitz der Erde doch im Laufe von Milliarden von Jahren immer wieder verandert. Und uberall trifft man auf die Spuren dieser unruhigen Vergangenheit. Richard Fortey ist ihr Chronist, und indem er uns an geologisch besonders interessante Statten fuhrt und deren Eigenheiten erlautert, lehrt er uns, das Wesen der Erde besser zu verstehen. Er zeigt, dass nicht nur die Gestalt der Erdoberflache, sondern auch die menschliche Kultur, die Naturgeschichte, ja sogar die Form unserer Stadte auf tieferen geologischen Prozessen beruhen. Die Reise beginnt an den Hangen des Vesuvs, wo Fortey die Geschichte dieser von Vulkanausbruchen gekennzeichneten Landschaft durch die Augen der Italiener des 15. Jahrhunderts, der Romer und - auf der Basis einzigartiger geologischer Befunde - der Menschen der Jungsteinzeit erzahlt. Und mit jeder neuen Geschichte, die er erzahlt, treten Verbindungen von der jungeren Vergangenheit zu langst vergessenen Zeiten zutage - bis tief hinab zu fernen geologischen Epochen, wenn er Plattenverschiebungen und die Bildung von alten Kontinenten und Meeren beschreibt. Nichts in diesem Buch scheint still zu stehen. Die Erdoberflache weitet sich und zieht sich wieder zusammen, Berge und Seen entstehen und vergehen, Kontinente driften umher und kollidieren. Unter Forteys kundiger Fuhrung erklimmen wir die Alpen, baden in den heissen Quellen Islands und tauchen hinab zum Meeresgrund. Wir erkunden die kahlen Felsen von Neufundland, klettern in bohmische Silberminen hinab, spazieren durch die uppigen Okosysteme von Hawaii, durchqueren die Salzebenen von Oman und schlendern am Andreas-Graben entlang. Forteys Beschreibungen der Schonheiten der Natur sind dabei so unvergesslich wie die besten Reiseberichte, seine Prosa ist so packend wie die eines Romanciers, und seine kristallklaren wissenschaftlichen Erklarungen sind faszinierend und oft uberraschend. Dieses wahrhaft tiefschurfende Buch wird Ihren Blick auf die Welt verandern - fur immer."

"Trilobites are the most lifelike of fossils many well-preserved specimens belie their great antiquity and seem almost ready to arch their bodies, peer about with their compound eyes, and crawl forward as if to complete a journey that was interrupted hundreds of millions of years ago." from the Foreword"New York State is and has long been a magnet for trilobite hunters. . . . New York's trilobites were among the first illustrated fossils in North America. . . . Many outstanding localities in New York State, from the majestic Ordovician limestone bluffs of Trenton Falls, to the Silurian beds in the great gorge of Niagara River, to the Devonian shale cliffs of Lake Erie, continue to yield abundant and spectacular trilobite fossils. New York strata have also yielded more trilobites with preserved appendages and other "soft parts" than almost any other region of the world. . . . Spectacular, ornate trilobites from New York ranging from a few millimeters to nearly a half meter in length, are featured in museums all over the world." from the PrefaceThis superbly illustrated book reviews the trilobite fossils found throughout New York State, including their biology, methods of taphonomy (preservation of specimens), and the broader Paleozoic geology of the state. A general chapter on the geology of New York State places the importance of these now-extinct invertebrate marine animals into context. Sixty-seven line drawings and 175 black-and-white photographs illustrate individual species, many represented here by type specimens, and display the eerie beauty that has made New York State trilobites favorites of collectors the world over."

Le genre Arvicola est considere comme un outil biostratigraphique majeur pour l'etude du Quaternaire. Les determinations specifiques d'apres l'indice d'email, SDQ, ont identifie plusieurs chronoespeces au sein de l'espece Arvicola cantiana. Cette etude revise le genre Arvicola en terme de morphodiversite et d'espaces morphologiques en utilisant l'analyse de contour et la biometrie. Ces methodes suggerent qu'une seule espece fossile, Arvicola cantiana, etait presente durant le Pleistocene. La grande tolerance climatique de cette espece lui permet d'etre repandue durant le Pleistocene moyen et le Pleistocene superieur, pendant les deux periodes glaciaire et interglaciaire. Notre etude expose la grande plasticite phenotypique permettant a l'espece de resister a des changements majeurs de l'environnement. Ces resultats remettent en question la fiabilite du SDQ pour les determinations specifiques et jette un doute sur l'elaboration d'une structure biochronologique basee sur Arvicola.