The emphasis in this volume is on the structure and functional design of the integument. The book starts with a brief introduction to some basic principles of physics (mechanics) including Newton’s Three Laws of Motion. These principles are subsequently used to interpret the problems animals encounter in motion. It is in only the last 40 or so years that we have begun to understand how important a role the integument plays in the locomotion of many marine vertebrates.  This involves the crossed-fiber architecture, which was first discovered in a classic study on nemertean worms. As a design principle we see that the crossed-fiber architecture is ubiquitous in nature. Research on some of the most dynamic marine vertebrates of the oceans – tuna, dolphins and sharks, and the extinct Jurassic ichthyosaurs – shows precisely how the crossed-fiber architecture contributes to high-speed swimming  and (in lamnid sharks) may even aid in energy conservation.  However, this design principle is not restricted to animals in the marine biota but is also found as far afield as the dinosaurs and, most recently, has been revealed as a major part of the microstructure of the most complex derivative of the integument, the feather. We see that a variety of phylogenetically diverse vertebrates take to the air by using skin flaps to glide from tree to tree or to the ground, and present detailed descriptions of innovations developed in pursuit of improved gliding capabilities in both extinct and modern day gliders. But the vertebrate integument had even greater things in store, namely true or flapping flight. Pterosaurs were the first vertebrates to use the integument as a membrane in true flapping flight and these interesting extinct animals are discussed on the basis of past and cutting-edge research , most intriguingly with respect to the structure of the flight membrane. Bats, the only mammals that fly, also employ integumental flight membranes. Classic research on bat flight is reviewed and supplemented with the latest research, which shows the complexities of the wing beat cycle to be significantly different from that of birds, as revealed by particle image velocimetry. The book’s largest chapter is devoted to birds, given that they make up nearly half of the over 22,000 species of tetrapods. The flight apparatus of birds is unique in nature and is described in great detail, with innovative research highlighting the complexity of the flight structures, bird flight patterns, and behavior in a variety of species. This is complimented by new research on the brains of birds, which shows that they are more complex than previously thought. The feather made bird flight possible, and was itself made possible by β-keratin, contributing to what may be a unique biomechanical microstructure in nature, a topic discussed in some depth. A highly polarized subject concerns the origin of birds and of the feather. Alleged fossilized protofeathers (primal simple feathers) are considered on the basis of histological and taphonomic investigative studies in Chapter 6. Finally, in Chapter 7 we discuss the controversies associated with this field of research. Professor Theagarten Lingham-Soliar works at the Nelson Mandela Metropolitan University, Port Elizabeth and is an Honorary Professor of Life Sciences at the University of KwaZulu-Natal.

The vertebrate integument arose about 450 million years ago as an 'armour' of dermal bony plates in small, jawless fish-like creatures, informally known as the ostracoderms. This book reviews the major changes that have occurred in the vertebrate integument from its beginnings to the present day. Critical questions concerning the origin, structure and functional biology of the bony integument are discussed and intrinsically linked to major steps in vertebrate evolution and phylogeny-the origin of jaws and the origin of teeth. The discussions include the origins of mineralization of major vertebrate skeletal components such as the dermatocranium, branchial arches and vertebral column. The advances that led to the origin of modern fishes and their phylogenetic development are reviewed and include the evolution of fins and replacement of the bony plates with several types of dermal scales. The evolution of reptiles saw a major transformation of the integument, with the epidermis becoming the protective outermost layer, from which the scales arose, while the dermis lay below it. The biological significance of the newly-evolved -keratin in reptilian scales, among the toughest natural materials known, is discussed in the context of its major contribution to the great success of reptiles and to the evolution of feathers and avian flight. The dermis in many vertebrates is strengthened by layers of oppositely oriented cross-fibres, now firmly entrenched as a design principle of biomechanics. Throughout the book conventional ideas are discussed and a number of new hypotheses are presented in light of the latest developments. The long evolutionary history of vertebrates indicates that the significance of the Darwinian concept of "survival of the fittest" may be overstated, including in our own mammalian origins and that chance often plays a major role in evolutionary patterns. Extensive illustrations are included to support the verbal descriptions. Professor Theagarten Lingham-Soliar is in the Department of Life Sciences at the University of KwaZulu-Natal.

This book addresses the principles and methods for determining petroleum source rocks based on fossil spores and pollen. Studying petroliferous basins in China, we discovered that there are often as many as three different sources of the microfossils: the source rocks, the rocks along the pathway, and the reservoir rocks. Therefore, fossil spores, pollen and algae from inland and coastal shelf petroliferous basins are analyzed and illustrated to show this complex process. Furthermore, the organic origin theory of oil is proven and environmental characteristics for hydrocarbon source-rock formation are discussed. Along with the geochronical and geographic distribution of non-marine petroleum source rocks in China, the mechanisms of petroleum migration following the pathways to the reservoirs are investigated. It will be a valuable reference work as well as a textbook for a wider research areas ranging from stratigraphy, palynology, palaeontology and petroleum geology.

Here, a diverse group of geologists and paleobiologists focus their attention on the richly fossiliferous Neogene stratigraphic sections of the Dominican Republic. They provide an updated geological framework and a series of novel studies of evolutionary stasis and change among different lineages and associated ecological communities. This collection of studies illustrates the immense potential of collaborative, multidisciplinary, and field-based paleobiological research.

This book presents the multidisciplinary results of an extensive underwater excavation in north Florida. This yielded the most complete results of interactions between early Paleoindians and late Pleistocene megafauna, in a rich environmental context in eastern North America. The data provides fundamental insights into "the Peopling of the Americas" and "The Extinction of the Megafauna". An excellent color photo section expresses the uniqueness of this project.

South American ecosystems suffered one of the greatest biogeographical events, after the establishment of the Panamian land bridge, called the "Great American Biotic Interchange" (GABI). This refers to the exchange, in several phases, of land mammals between the Americas; this event started during the late Miocene with the appearance of the Holartic Procyonidae (Huayquerian Age) in South America and continues today. The major phases of mammalian dispersal occurred from the Latest Pliocene (Marplatan Age) to the Late Pleistocene (Lujanian Age). The most important and richest localities of Late Miocene-Holocene fossil vertebrates of South America are those of the Pampean region of Argentina. There are also several Late Miocene and Pliocene localities in western Argentina and Bolivia. Other important fossils have been collected in localities of Pleistocene age outside Argentina: Tarija (Bolivia), karstic caves of Lagoa Santa and the recently explored caves of Tocantins (Brasil), Talara (Peru), La Carolina (Ecuador), Muaco (Venezuela), and Cueva del Milodon (Chile), among others. The book discusses basic information for interpreting the GABI such as taxonomic composition (incorporating the latest revisions) at classical and new localities for each stage addressing climate, environments, and time boundaries for each stage. It includes the chronology and dynamics of the GABI, the integration of South American mammalian faunas through time, the Quaternary mammalian extinctions and the composition of recent mammalian fauna of the continent.

This book gathers and updates the most significant advances of the last two centuries and presents an unprecedented micro paleontological study covering more than 20 stratigraphic sections. This information is supplemented by numerous sedimentological observations and analyses, on the basis of which a new lithostratigraphic framework for the Neogene of the Chacoparanense Basin is proposed. The book is structured in an easy-to-read format: Its main section offers a comprehensive review of the current state of knowledge on transgressions in Argentina and similar transgressions in other South American countries, taking into account various key aspects (age, paleoenvironment, micropaleontology, etc.). Secondly, the book presents the main results on the TLP and TEP of the Chacoparanense Basin and the TEP of the Peninsula de Valdes. Lastly, it provides readers with complete stratigraphic profiles (Appendix A), mineralogical analyses (Appendix B), distribution charts (Appendix C), systematics (Appendix D) and plates (Appendix E).

The Late Cenozoic Yushe Basin, Shanxi Province, China embodies the bulk of our knowledge on successions of terrestrial vertebrates in the northern part of East Asia. Everything we know about Asian mammals of the last 6 million years has a historical basis in the documentation of the geology of Yushe. This volume introduces the basin in its geological setting, describes the succession of fossiliferous strata, and shows how it is dated. It develops an unsurpassed level of precision for its age control. Geological maps and stratigraphic sections provide the backbone for individual studies to follow on varied fossil groups. The volume explores the history of exploration of the last century in Yushe Basin and places development of paleontology there into the context of the birth of the modern epoch of science in China.

Carbonate sediments are of increasing relevance for archives of past environmental conditions and for economical reasons in areas of geothermal energy and hydrocarbon reservoirs. Complex interaction of physical and chemical parameters with biological parameters determines the architecture and composition of carbonate sedimentary bodies. This book closes some of the still existing gaps in our understanding of the influence and interplay of physical, chemical, and biological parameters with carbonate sedimentation. An understanding of this interaction is not only required for reliable prediction of reservoir quality but also for a robust interpretation of environmental conditions in the past and the present. It is written by geologists for geologists in order to provide an easily accessible overview of the large amount of relevant information provided by the neighbouring sciences. The approach of the book is to document the modern depositional environments of three classical areas of carbonate deposition, each characteristic for a specific sedimentological setting (isolated platform, attached shelf, ramp) in order to assess both the range of physical, biological and chemical parameters and their sedimentary response. This book presents a comprehensive compilation based on data from published work and unpublished theses, and the integration of these data in order to extract previously undiscovered relationships between the discussed parameters and carbonate deposition.

Two of the greatest evolutionary events in the history of life on Earth occurred during Early Paleozoic time. The first was the Cambrian explosion of skeletonized marine animals about 540 million years ago. The second was the "Great Ordovician Biodiversification Event," which is the focus of this book. During the 46-million-year Ordovician Period (489--443 m.y.), a bewildering array of adaptive radiations of "Paleozoic- and Modern-type" biotas appeared in marine habitats, the first animals (arthropods) walked on land, and the first non-vascular bryophyte-like plants (based on their cryptospore record) colonized terrestrial areas with damp environments.

This book represents a compilation by a large team of Ordovician specialists from around the world, who have enthusiastically cooperated to produce this first globally orientated, internationally sponsored IGCP (International Geological Correlation Program) project on Ordovician biotas. The major part is an assembly of genus- and species-level diversity data for the many Ordovician fossil groups. The book also presents an evaluation of how each group diversified through Ordovician time, with assessments of patterns of change and rates of origination and extinction. As such, it will become the standard work and data source for biotic studies on the Ordovician Period.

This book serves as an introduction to the Cretaceous geology and palaeontology of south-central Africa, covering the whole of Southern and Eastern Africa and Angola. Fifty two plates illustrate almost 1000 species and provide a field guide to the macrofossils of the subcontinent. The book will be of value to field geologists, students and non-specialists with an interest in the natural world. A bibliography of the Cretaceous palaeontology and stratigraphy of the subcontinent is provided. Features: Provides a concise account of the Cretaceous geology for 13 African regions Includes beautiful illustrations and a comprehensive bibliography Fossils are presented in stratigraphical order, allowing easy determination of the age deposits.

This book presents a detailed examination of the current state of knowledge in the field of paleoneurology in the main amniote groups (reptiles, birds and mammals), and advances resulting from new non-invasive technologies. The study of fossil endocasts is an area of considerable current interest, and has long been central to our understanding of the evolution of the brain, development of senses and behavioral adaptations in diverse vertebrate groups and across vertebrates as a whole. Recent advances in non-invasive imaging have significantly increased the number of fossil taxa for which brain morphology is known, and it may now be possible to quantitatively analyze the relative size of brain regions. Providing a general overview of current perspectives and problems in evolutionary neuroanatomy, this book is intended for a wide range of readers, including undergraduate and graduate students, teachers, and anyone with a special interest in paleoneurology. It is also useful as supplementary reading for courses in digital anatomy, vertebrate comparative anatomy, computed morphometrics, paleontology, neurology and radiology as well as evolution programs

We cannot catechise our stony ichthyolites, as did the necromantic lady of the Arabian Nights did the coloured fishes of the lake which had once been a city, when she touched their dead bodies with her wand, and they straightaway raised their heads and rephed to her queries. We would have many a question to ask them if we could - questions never to be solved. Hugh Miller, The Old Red Sandstone When I started this book in 1991, the subject of vertebrate origins was fusty and unfashionable. Early drafts for this preface read like an extend ed complaint at the lot of traditional morphologists, cast aside by the march of modern molecular biology. But no longer - this book should reach you at a time of renewed inter est in the origin of the vertebrates, our own particular corner of creation. For although the topic has excited interest for well over a century, molec ular biology has only lately achieved the maturity necessary to test its predictions. As a legitimate field of study, it is fashionable again."

The climate of the Earth has undergone many changes and for those times when geologic data are widespread and abundant the Mesozoic appears to have been one of the warmest intervals. This was a time during which the single continent Pangea disintegrated into continental units similar to those of today, a time when there were no significant polar ice caps and sea level was generally much higher than at the present time, and a time when dinosaurs apparently dominated terrestrial faunas and the flowering plants evolved. Understanding this alien world, ancestral to ours, is intrinsically interesting, intellectually challenging, and offers opportunities for more effective targeting of sites where commercially important geological resources may be found. It also provides critical insights into the operation of coupled Earth systems (biospheric, atmospheric, hydrospheric and geospheric) under extreme 'greenhouse' conditions, and therefore may have relevance to possible future global change. Our intention in organizing this Discussion Meeting was to bring together those who gather and interpret geologic data with those who model global climates from first principles. The community of workers who study the Quaternary have made significant advances by integrating and comparing palaeodata and climate model experiments. Although we have focused not on the Quaternary 'icehouse' but on the Mesozoic 'hothouse' climate we are well aware that approaches used in the study of the Quaternary may have relevance to earlier times.

Multispecies Archaeology explores the issue of ecological and cultural novelty in the archaeological record from a multispecies perspective. Human exceptionalism and our place in nature have long been topics of academic consideration and archaeology has been synonymous with an axclusively human past, to the detriment of gaining a more nuanced understanding of one that is shared. Encompassing more than just our relationships with animals, the book considers what we can learn about the human past without humans as the focus of the question. The volume digs deep into our understanding of interaction with plants, fungi, microbes, and even the fundamental building blocks of life, DNA. Multispecies Archaeology examines what it means to be human-and non-human-from a variety of perspectives, providing a new lens through which to view the past. Challenging not only the subject or object of archaeology but also broader disciplinary identities, the volume is a landmark in this new and evolving area of scholarly interest.

ANOXIA defines the lack of free molecular oxygen in an environment. In the presence of organic matter, anaerobic prokaryotes produce compounds such as free radicals, hydrogen sulfide, or methane that are typically toxic to aerobes. The concomitance of suppressed respiration and presence of toxic substances suggests these habitats are inhospitable to Eukaryota. Ecologists sometimes term such environments 'Death Zones'. This book presents, however, a collection of remarkable adaptations to anoxia, observed in Eukaryotes such as protists, animals, plants and fungi. Case studies provide evidence for controlled beneficial use of anoxia by, for example, modification of free radicals, use of alternative electron donors for anaerobic metabolic pathways, and employment of anaerobic symbionts. The complex, interwoven existence of oxic and anoxic conditions in space and time is also highlighted as is the idea that eukaryotic inhabitation of anoxic habitats was established early in Earth history.

This volume brings together a series of papers that address the topic of reconstructing behavior in the primate fossil record. The literature devoted to reconstructing behavior in extinct species is ovelWhelming and very diverse. Sometimes, it seems as though behavioral reconstruction is done as an afterthought in the discussion section of papers, relegated to the status of informed speculation. But recent years have seen an explosion in studies of adaptation, functional anatomy, comparative sociobiology, and development. Powerful new comparative methods are now available on the internet. At the same time, we face a rapidly growing fossil record that offers more and more information on the morphology and paleoenvironments of extinct species. Consequently, inferences of behavior in extinct species have become better grounded in comparative studies of living species and are becoming increas ingly rigorous. We offer here a series of papers that review broad issues related to reconstructing various aspects of behavior from very different types of evi dence. We hope that in so doing, the reader will gain a perspective on the various types of evidence that can be brought to bear on reconstructing behavior, the strengths and weaknesses of different approaches, and, perhaps, new approaches to the topic. We define behavior as broadly as we can including life-history traits, locomotion, diet, and social behavior, giving the authors considerable freedom in choosing what, exactly, they wish to explore."

The Qattara Depression is part of the Northwestern Desert in Egypt and is home to the second lowest point in Africa at -133 meters below sea level. Therefore, before any projects can be carried out in this area, we must first understand the geology of the land. The present study deals with the high-resolution sequence stratigraphic analysis of the Lower Miocene Moghra Formation outcrops in the Qattara Depression Region. The literature on the sedimentology and sequence stratigraphy of the Moghra Formation has been sparse to date, despite some excellent work over the years by academic and petroleum workers. Moreover, the area studied is within what was once a front-line of World War II, where mine fields and war relics are scattered and cover wide reaches. This has resulted in limited geologic mapping in the past. Thus, great attention is paid in this study to establishing a robust sedimentology and high-resolution sequence stratigraphic framework for the Lower Miocene Moghra Formation. Included are works based on outcrops and, most importantly, new sedimentological and chronostratigraphic information not previously available.

"[An] extraordinary book. . . . Mr. Gould is an exceptional combination of scientist and science writer. . . . He is thus exceptionally well placed to tell these stories, and he tells them with fervor and intelligence."—James Gleick, New York Times Book Review

High in the Canadian Rockies is a small limestone quarry formed 530 million years ago called the Burgess Shale. It hold the remains of an ancient sea where dozens of strange creatures lived—a forgotten corner of evolution preserved in awesome detail. In this book Stephen Jay Gould explores what the Burgess Shale tells us about evolution and the nature of history.

"Gould at his best. . . . The message of history is superbly conveyed. . . . Recommended reading for scientists and nonscientists of all persuasions."—Walter C. Sweet, Science

"Luminous. . . . Filled with profound and upsetting ideas like the Burgess Shale itself and just as solid. It is surely one of nature's best stories, told with a light touch by a master of the field."—Lewis Thomas, M.D.

"There is no question about the historical importance of the Burgess Shale, and Gould is right when he says that it deserves a place in the public consciousness along with big bangs and black holes. . . . A compelling story, told with characteristic verve."—Richard A. Fortey, Nature

Fossil species appear to persist morphologically unchanged for long intervals of geologic time, punctuated by short bursts of rapid change as explained by the Ecological Evolutionary Units (EEUs). Here, morphological variation in Paleozoic atrypide morphology at the subfamily level (Atrypinae and Variatrypinae) from the Silurian and Devonian time intervals in the third Paleozoic EEU (~444-359 my) were investigated using relatively new techniques of quantitative modeling. The study explains how a group of closely related taxa in atrypide subfamilies exhibit morphological conservation through time in P3 EEU within the Eastern North America region.

The study ofanthropoid origins continues to be a lightning rod for research in paleoanthropology. Issuessurrounding anthropoid origins impact the higher leveltaxonomy ofprimates, adaptivescenariosfor basalprimate radiations, and the timing of origination of the major primate clades. Basic questions about anthropoid evolution remain unanswered. Where do anthropoids fit phyloge- netically among primates? Where and when did the group originate? What functional and adaptive innovations characterize anthropoids today and what is the adaptive significanceand phylogenetic history ofthese innovations? The fossil record of early anthropoid evolution has greatly improved in recent years. Developments in systematictechniques and theory, as well as the burgeoning molecular evidence, make this an ideal time for these fossil discoveries to be placed in the context of data on the relationships among living primates. There isan improved understandingoffunction and adaptation in the visual system, brain, and masticatory apparatus, key anatomical systems where anthropoid synapomorphies are concentrated. New methods for estimating visualacuity and activitypatterns in fossil primates are providing insights into the evolution ofthe visualsystem. The rapid accumulation ofinformation on color vision in primates, including new genetic evidence of possible trichro- macyin strepsirrhines, and new behavioraldata on the benefitsofcolor vision, makes this an exciting time to evaluate the role of chromatic perception in anthropoid evolution. Research into the primate visualsystem by neuroscien- tists has generated a plethoraofimportant data in recent years, making this an ideal time to bring these researchers together with anthropologists.

This book covers the hot topics of angiosperm structure and evolution in several chapters discussing vegetative and reproductive characters. It also looks at the implications of ancestral angiosperm characters for an herbaceous origin and the phylogeny of angiosperms from a structure and molecular perspective.

Relying on the latest analytical techniques, this all-embracing new reference offers comprehensive coverage of the development, evolution, and morphology of both fossil and living cephalopods. In 34 in-depth chapters a group of 51 international neontologists and peleontologists offer and opverview of current methods, data, analyses, and interpretations, and posit suggestions for future research. With its unparalleled combination of first-rate contributions on living and fossil cephalopods, this book provides researchers and advanced students in paleontology, invertebrate zoology, evolutionary biology, and allied disciplines with a trove of recent data and authoritative interpretations that will immeasurably benefit their own studies.

The ocean is the womb of all life on earth. It is beautiful and bizarre, violent and mysterious. Inhabited by a cast of characters stolen from fantasy, it's a dystopian world where dragons are real, and monsters are commonplace. Today's spectacular marine life has an ancient history preserved in stone - fossil strata that read like dramatic pages from the longest story ever told - tales of evolution, extinction, and surprising continuity. Having thrived a tumultuous 500 million years, this marine kingdom is now challenged by a new, arrogant and domineering life form. This book looks beyond the media focus on climate change and extinction to celebrate the continuity of ocean life. I'll take you on a personal journey to explore origins and destinies, from primordial soup to today's threatened oceans - towards a future we can influence. We always have a choice.

This volume brings together a series of papers that address the topic of reconstructing behavior in the primate fossil record. The literature devoted to reconstructing behavior in extinct species is ovelWhelming and very diverse. Sometimes, it seems as though behavioral reconstruction is done as an afterthought in the discussion section of papers, relegated to the status of informed speculation. But recent years have seen an explosion in studies of adaptation, functional anatomy, comparative sociobiology, and development. Powerful new comparative methods are now available on the internet. At the same time, we face a rapidly growing fossil record that offers more and more information on the morphology and paleoenvironments of extinct species. Consequently, inferences of behavior in extinct species have become better grounded in comparative studies of living species and are becoming increas ingly rigorous. We offer here a series of papers that review broad issues related to reconstructing various aspects of behavior from very different types of evi dence. We hope that in so doing, the reader will gain a perspective on the various types of evidence that can be brought to bear on reconstructing behavior, the strengths and weaknesses of different approaches, and, perhaps, new approaches to the topic. We define behavior as broadly as we can including life-history traits, locomotion, diet, and social behavior, giving the authors considerable freedom in choosing what, exactly, they wish to explore."