Research on learning and cognition in geoscience education research and other discipline-based education communities suggests that effective instruction should include three key components: a) activation of students' prior knowledge on the subject, b) an active learning pedagogy that allows students to address any existing misconceptions and then build a new understanding of the concept, and c) metacognitive reflections that require students to evaluate their own learning processes during the lesson. This Element provides an overview of the research on student-centered pedagogy in introductory geoscience and paleontology courses and gives examples of these instructional approaches. Student-centered learning shifts the power and attention in a classroom from the instructor to the students. In a student-centered classroom, students are in control of their learning experience and the instructor functions primarily as a guide. Student-centered classrooms trade traditional lecture for conceptually-oriented tasks, collaborative learning activities, new technology, inquiry-based learning, and metacognitive reflection.

Hands-on learning in paleontology, and geology in general, is fairly common practice. Students regularly use rocks, fossils, and data in the classroom throughout their undergraduate career, but they typically do it sitting in a chair in a lab. Kinesthetic learning is a teaching model that requires students to be physically active while learning. Students may be involved in a physical activity during class or might be using their own bodies to model some important concept. This Element briefly discusses the theory behind kinesthetic learning and how it fits into a student-centered, active-learning classroom. It then describes in detail methods for incorporating it into student exercises on biostratigraphy, assessment of sampling completeness, and modeling evolutionary processes. Assessment data demonstrates that these exercises have led to significantly improved student learning outcomes tied to these concepts.

Integration of research experiences into the undergraduate classroom can result in increased recruitment, retention, and motivation of science students. 'Big data' science initiatives, such as the Paleobiology Database (PBDB), can provide inexpensive and accessible research opportunities. This Element provides an introduction to what the PBDB is, how to use it, how it can be deployed in introductory and advanced courses, and examples of how it has been used in undergraduate research. The PBDB aims to provide information on all fossil organisms, across the tree of life, around the world, and through all of geologic time. The PBDB Resource Page contains a range of PBDB tutorials and activities for use in physical geology, historical geology, paleontology, sedimentology, and stratigraphy courses. As two-year colleges, universities, and distance-based learning initiatives seek research-based alternatives to traditional lab exercises, the PBDB can provide opportunities for hands-on science activities.

People hold a variety of prior conceptions that impact their learning. Prior conceptions that include erroneous or incomplete understandings represent a significant barrier to durable learning, as they are often difficult to change. While researchers have documented students' prior conceptions in many areas of geoscience, little is known about prior conceptions involving paleontology. In this Element, data on student prior conceptions from two introductory undergraduate paleontology courses are presented. In addition to more general misunderstandings about the nature of science, many students hold incorrect ideas about methods of historical geology, Earth history, ancient life, and evolution. Of special note are student perceptions of the limits of paleontology as scientific inquiry. By intentionally eliciting students' prior conceptions and implementing the pedagogical strategies described in other Elements in this series, lecturers can shape instruction to challenge this negative view of paleontology and improve student learning.

Lecturing has been a staple of university pedagogy, but a shift is ongoing because of evidence that active engagement with content helps strengthen learning and build more advanced skills. The flipped classroom, which delivers content to students outside of the class meeting, is one approach to maximize time for active learning. The fundamental benefit of a flipped class is that students learn more, but ensuring student preparation and engagement can be challenging. Evaluation policies can provide incentives to guide student effort. Flipping a class requires an initial time commitment, but the workload associated with evaluating student work during the course can be mitigated. The personal interactions from active learning are extremely rewarding for students and instructors, especially when class sizes are small and suitable room layouts are available. Overall, flipping a course doesn't require special training, just a willingness to experiment, reflect, and adjust.

Research-led, research-oriented, and research-based teaching incorporate research into teaching to different degrees. Research-led teaching focuses on content and informs students about current research findings, while research-oriented teaching focuses on techniques and often occurs in research methods courses. In research-based teaching, students participate in research. Through this involvement, they benefit from improved content knowledge, research skills, and life skills, as well as enhanced personal development. Research-embedded courses can make such benefits available to a wide range of students. Best practices in experiential learning and the incorporation of research in teaching include intentionality, planning, authenticity, reflection, training, monitoring, assessment, and acknowledgment. In this Element, these principles of best practice are illustrated by courses with embedded student research. Guidelines are presented for how to plan and execute a semester-long course-embedded research project, as well as alternative and shorter-term approaches. Research-based teaching provides challenges for students and faculty, but the benefits for all stakeholders are strong.

The educational benefits of replacing in-class lectures with hands-on activities are clear. Such active learning is a natural fit for paleontology, which can provide opportunities for examining fossils, analyzing data and writing. Additionally, there are a number of topics in the field that are exciting to geology majors and non-majors alike: very few can resist the lure of dinosaurs, huge meteor impacts, vicious Cretaceous sharks or a giant Pleistocene land mammal. However, it can seem difficult to introduce these techniques into a large general education class full of non-majors: paleontological specimens provide a natural starting point for hands-on classroom activities, but in a large class it is not always practical or possible to provide enough fossil material for all students. The Element introduces different types of active learning approaches, and then explains how they have been applied to a large introductory paleontology class for non-majors.

University dinosaur courses provide an influential venue for developing aptitude beyond knowledge of terrestrial Mesozoic reptiles. Passion for dinosaurs, when properly directed, can trigger interest in science and be used to develop critical thinking skills. Examination of dinosaur paleontology can develop competence in information analysis, perception of flawed arguments, recognition of persuasion techniques, and application of disciplined thought processes. Three methods for developing critical thought are outlined in this Element. The first uses dinosaur paleontology to illustrate logical fallacies and flawed arguments. The second is a method for evaluating primary dinosaur literature by students of any major. The final example entails critique of dinosaur documentaries based on the appearance of dinosaurs and the disconnect between scientific fact and storytelling techniques. Students are owed more than dinosaur facts; lecturers should foster a set of skills that equips students with the tools necessary to be perceptive citizens and science advocates.