How do radically new kinds of organisms evolve? The Origin of Higher Taxa addresses this essential question, specifically whether the emergence of higher taxa such as orders, classes, and phyla are the result of normal Darwinian evolution acting over a sufficiently long period of time, or whether unusual genetic events and particular environmental and ecological circumstances are also involved. Until very recently, the combination of an incomplete fossil record and a limited understanding about how raw mutations lead via modified ontogenic processes to significant phenotypic changes, effectively stymied scientific debate. However, it is now timely to revisit the question in the light of the discovery of considerable new fossil material (and new techniques for studying it), together with significant advances in our understanding of phenotypic development at the molecular level. This novel text incorporates evidence from morphology, palaeobiology, developmental biology, and ecology, to review those parts of the fossil record that illustrate something of the pattern of acquisition of derived characters in lineages leading to actual higher taxa as well as the environmental conditions under which they occurred. The author's original ideas are set within the context of a broad and balanced review of the latest research in the field. The result is a book which provides a concise, authoritative, and accessible overview of this fascinating subject for both students and researchers in evolutionary biology and palaeontology.

Codon-based models of evolution are a relatively new addition to the toolkit of computational biologists, and in recent years remarkable progress has been made in this area. The study of evolution at the codon level captures information contained in both amino acid and synonymous DNA substitutions. By combining these two types of information, codon analyses are more powerful than those of either amino acid or DNA evolution alone. This is a clear benefit for most evolutionary analyses, including phylogenetic reconstruction, detection of selection, ancestral sequence reconstruction, and alignment of coding DNA. Despite the theoretical advantages of codon based models, their relative complexity delayed their widespread use. Only in recent years, when large-scale sequencing projects produced sufficient genomic data and computational power increased, did their usage become more common. In Codon Evolution, leading researchers in the field of molecular evolution provide the latest insights from codon-based analyses of genetic sequences. The first part of the book provides comprehensive coverage of the developments of various types of codon substitution models such as parametric and empirical models used in maximum likelihood as well as Bayesian frameworks. Subsequent chapters examine the use of codon models to infer selection and other applications of codon models to biological systems. The second part of the book focuses on codon usage bias. Both the underlying mechanisms as well as current methods to analyse codon usage bias are presented.

Animal Evolution provides a comprehensive analysis of the evolutionary interrelationships and myriad diversity of the Animal Kingdom. It reviews the classical, morphological information from structure and embryology, as well as the new data gained from studies using immune stainings of nerves and muscles and blastomere markings which makes it possible to follow the fate of single blastomeres all the way to early organogenesis. Until recently, the information from analyses of gene sequences has tended to produce myriads of quite diverging trees. However, the latest generation of molecular methods, using many genes, expressed sequence tags, and even whole genomes, has brought a new stability to the field. For the first time this book brings together the information from these varied fields, and demonstrates that it is indeed now possible to build a phylogenetic tree from a combination of both morphology and gene sequences. This thoroughly revised third edition of Animal Evolution brings the subject fully up to date, especially in light of the latest advances in molecular techniques. The book is generously illustrated throughout with finely detailed line drawings and clear diagrams, many of them new.

Primary sexual traits, those structures and processes directly involved in reproduction, are some of the most diverse, specialized, and bizarre in the animal kingdom. Moreover, reproductive traits are often species-specific, suggesting that they evolved very rapidly. This diversity, long the province of taxonomists, has recently attracted broader interest from evolutionary biologists, especially those interested in sexual selection and the evolution of reproductive strategies.
Primary sexual characters were long assumed to be the product of natural selection, exclusively. A recent alternative suggests that sexual selection explains much of the diversity of "primary" sexual characters. A third approach to the evolution of reproductive interactions after copulation or insemination has been to consider the process one of sexual conflict. That is, the reproductive processes of a species may reflect, as does the mating system, evolution acting on males and on females, but in different directions.
In this volume, authors explore a wide variety of primary sexual characters and selective pressures that have shaped them, from natural selection for offspring survival to species-isolating mechanisms, sperm competition, cryptic female choice and sexual arms races. Exploring diverse reproductive adaptations from a theoretical and practical perspective, The Evolution of Primary Sexual Characters will provide an unparalleled overview of sexual diversity in many taxa and an introduction to the issues in sexual selection that are changing our view of sexual processes.

Animal life, now and over the past half billion years, is incredibly diverse. Describing and understanding the evolution of this diversity of body plans - from vertebrates such as humans and fish to the numerous invertebrate groups including sponges, insects, molluscs, and the many groups of worms - is a major goal of evolutionary biology. In this book, a group of leading researchers adopt a modern, integrated approach to describe how current molecular genetic techniques and disciplines as diverse as palaeontology, embryology, and genomics have been combined, resulting in a dramatic renaissance in the study of animal evolution.
The last decade has seen growing interest in evolutionary biology fuelled by a wealth of data from molecular biology. Modern phylogenies integrating evidence from molecules, embryological data, and morphology of living and fossil taxa provide a wide consensus of the major branching patterns of the tree of life; moreover, the links between phenotype and genotype are increasingly well understood. This has resulted in a reliable tree of relationships that has been widely accepted and has spawned numerous new and exciting questions that require a reassessment of the origins and radiation of animal life. The focus of this volume is at the level of major animal groups, the morphological innovations that define them, and the mechanisms of change to their embryology that have resulted in their evolution. Current research themes and future prospects are highlighted including phylogeny reconstruction, comparative developmental biology, the value of different sources of data and the importance of fossils, homology assessment, character evolution, phylogeny of major groups of animals, and genome evolution. These topics are integrated in the light of a 'new animal phylogeny', to provide fresh insights into the patterns and processes of animal evolution.
Animal Evolution provides a timely and comprehensive statement of progress in the field for academic researchers requiring an authoritative, balanced and up-to-date overview of the topic. It is also intended for both upper level undergraduate and graduate students taking courses in animal evolution, molecular phylogenetics, evo-devo, comparative genomics and associated disciplines.

This second edition of Field Guide to the Wild Flowers of the Western Mediterranean is the most comprehensive and up-to-date plant identification guide to the region. Encompassing an area from southern Europe to North Africa, the western Mediterranean is united by a climate of hot, dry summers and mild, damp winters. This region is famous for its biodiversity and has an exceptionally rich flora.

Animal phylogeny is undergoing a major revolution due to the availability of an exponentially increasing amount of molecular data and the application of novel methods of phylogentic reconstruction, as well as the many spectacular advances in palaeontology and molecular developmental biology. Traditional views of the relationships among major phyla have been shaken and new, often unexpected, relationships are now being considered. At the same tiem, the emerging discipline of evolutionary developmental biology, or 'evo-devo', has offered new insights into the origin and evolvability of major traits of animal architecture and life cycle. All these developments call for a revised interpretation of the pathways along which animal structure and development has evolved since the origin of the Metazoa.
Perspectives in Animal Phylogeny and Evolution takes on this challenge, successfully integrating morphological, fossil and molecular evidence to produce a novel reinterpretation of animal evolution. Central to the book's approach is an 'evo-devo' perspective on animal evolution (with all the fresh insights this has given into the origin of animal organization and life cycles), complementary to the more traditional perspectives of pattern (cladistics, comparative anatomy and embryology), mechanisms (developmental biology) and adaptation (evolutionary biology). The author advocates the need to approach the study of animal evolution with a critical attitude towards many key concepts of comparative morphology and developmental biology. Particular attention in the book is paid to the evolution of life cycles and larval forms.
This accessible text is suitable for graduate students takingadvanced courses in evolutionary developmental biology, invertebrate zoology, molecular phylogenetics and palaeontology, as well as professional researchers in these fields requiring an authoritative and up-to-date overview of this dynamic topic.

Even before the publication of Darwin's Origin of Species, the perception of evolutionary change has been a tree-like pattern of diversification - with divergent branches spreading further and further from the trunk. In the only illustration of Darwin's treatise, branches large and small never reconnect. However, it is now evident that this view does not adequately encompass the richness of evolutionary pattern and process. Instead, the evolution of species from microbes to mammals builds like a web that crosses and re-crosses through genetic exchange, even as it grows outward from a point of origin. Some of the avenues for genetic exchange, for example introgression through sexual recombination versus lateral gene transfer mediated by transposable elements, are based on definably different molecular mechanisms. However, even such widely different genetic processes may result in similar effects on adaptations (either new or transferred), genome evolution, population genetics, and the evolutionary/ecological trajectory of organisms. For example, the evolution of novel adaptations (resulting from lateral gene transfer) leading to the flea-borne, deadly, causative agent of plague from a rarely-fatal, orally-transmitted, bacterial species is quite similar to the adaptations accrued from natural hybridization between annual sunflower species resulting in the formation of several new species. Thus, more and more data indicate that evolution has resulted in lineages consisting of mosaics of genes derived from different ancestors. It is therefore becoming increasingly clear that the tree is an inadequate metaphor of evolutionary change. In this book, Arnold promotes the 'web-of-life' metaphoras a more appropriate representation of evolutionary change in all lifeforms.

Systematics has developed rapidly during the past two decades. A multitude of new methods and contributions from a diversity of biological fields including molecular genetics and developmental biology have provided a wealth of phylogenetic hypotheses, some confirming traditional views others contradicting them. Despite such inconsistencies, it is now possible to recognize robust regions of a 'tree of life' and also to identify problematic areas which have yet to be resolved. This is the first book to apply the current state of phylogeny to an evolutionary interpretation of animal organ systems and body architecture, providing alternative theories in those cases of continuing controversy.
Organs do not appear suddenly during evolution; instead they are composed of far simpler structures. In some cases it is even possible to trace particular molecules or physiological pathways as far back as pre-animal history. What emerges is a fascinating picture, showing how animals have combined ancestral and new elements in novel ways to form constantly changing responses to environmental requirements.
The Evolution of Organ Systems starts with a general overview of current animal phylogeny, followed by review of general body organization including symmetry, anteroposterior axis, dorsoventral axis, germ layers, segmentation, and skeletons. Subsequent chapters then provide a detailed description of the individual organ systems themselves - integument, musculature, nervous system, sensory organs, body cavities, excretory system, circulatory system, respiratory system, intestinal system, gonads and gametes. Generously illustrated throughout, this accessible text is suitable for both upper levelundergraduate and graduate students taking courses in animal evolution, organogenesis, animal anatomy, zoology and systematics. It will also be a valuable reference tool for those professional researchers in these fields requiring an authoritative, balanced and up-to-date overview of the topic.

Ancestral sequence reconstruction is a technique of growing importance in molecular evolutionary biology and comparative genomics. As a powerful tool for testing evolutionary and ecological hypotheses, as well as uncovering the link between sequence and molecular phenotype, there are potential applications in a range of fields. Ancestral Sequence Reconstruction starts with a historical overview of the field, before discussing the potential applications in drug discovery and the pharmaceutical industry. This is followed by a section on computational methodology, which provides a detailed discussion of the available methods for reconstructing ancestral sequences (including their advantages, disadvantages, and potential pitfalls). Purely computational applications of the technique are then covered, including whole proteome reconstruction. Further chapters provide a detailed discussion on taking computationally reconstructed sequences and synthesizing them in the laboratory. The book concludes with a description of the scientific questions where experimental ancestral sequence reconstruction has been utilized to provide insights and inform future research. This research level text provides a first synthesis of the theories, methodologies and applications associated with ancestral sequence recognition, while simultaneously addressing many of the hot topics in the field. It will be of interest and use to both graduate students and researchers in the fields of molecular biology, molecular evolution, and evolutionary bioinformatics.

In On the Origin of Species (1859), Charles Darwin presented his evidence for evolution and natural selection as its mechanism. He drew upon his earliest data gathered during his voyage on the HMS Beagle, which included collecting mammalian fossils in South America clearly related to living forms, tracing the geographical distributions of living species across South America, and sampling the peculiar fauna of the geologically young Galapagos Archipelago that showed evident affinities to South American forms. By the end of the voyage, he came to the realization that instead of various centers of creation, species evolved in different regions throughout the world. However, except for some personal ponderings, he did not express this revelation explicitly in his notebooks until shortly after his return. Over the years, he collected more evidence supporting evolution, but his early work remained paramount: it became the first paragraph of On the Origin of Species and encompassed three separate chapters, as well as later appearing in his autobiography. Many discussions of Darwin's landmark book give scant attention to this wealth of evidence and today we still do not fully appreciate its significance in Darwin's thinking. In Origins of Darwin's Evolution, J. David Archibald explores this lapse. He also shows that Darwin's other early passion, geology, proved a more elusive corroboration of evolution. On the Origin of Species dedicated only one chapter to the rock and fossil record, as it appeared too incomplete for Darwin's evidentiary standards. Carefully retracing Darwin's gathering of evidence and the evolution of his thinking, Origins of Darwin's Evolution achieves a new understanding of how Darwin crafted his transformative theory.

This edited volume is provides an authoritative synthesis of knowledge about the history of life. All the major groups of organisms are treated, by the leading workers in their fields. With sections on: The Importance of Knowing the Tree of Life; The Origin and Radiation of Life on Earth; The Relationships of Green Plants; The Relationships of Fungi; and The Relationships of Animals. This book should prove indispensable for evolutionary biologists, taxonomists, ecologists interested in biodiversity, and as a baseline sourcebook for organismic biologists, botanists, and microbiologists. An essential reference in this fundamental area.

Over-two thirds of the U.S. population is allergic to poison oak, poison ivy, or a related plant. These and many other common plants in our homes, fields, and gardens are irritants that cause misery to many. But surprisingly, there has never been a general guide to help raise awareness of them--and to help avoid them. This new book reviews the history, occurrence, classification, toxicity, and health aspects of all the major allergenic and irritant species.

Cephalopods are fast-moving, voracious predators, and can change colour with breath-taking rapidity. They range from the giant squid, the world's largest marine invertebrate, to species of only 2 cm in length. Inhabitants of most seas of the world, they are found from the surface to great depths. Most cephalopods have short lives yet their efficiency in capturing and consuming prey ensures rapid growth. These animals possess highly-developed nervous systems, large brains, elaborate senses, complex behaviour and are capable of learning. Many of these features are described and illustrated with line drawings and photomicrographs.

This volume describes and illustrates the microscopic anatomy of the "aroids", the large, economically important plant family Araceae, and also the basal monocot family Acoraceae.

This fully updated edition explores conceptual as well as technical guidelines for plant taxonomists and geneticists, such as the increasing use of next-generation sequencing (NGS) technologies for numerous applications in plant taxonomy. The volume provides molecular approaches to be used within an "integrative taxonomy" framework, combining a range of nucleic acid and cytogenetic data together with other crucial information (taxonomy, morphology, anatomy, ecology, reproductive biology, biogeography, paleobotany, etc.), which will help not only to best circumvent species delimitation but also to resolve the evolutionary processes in play. Written for the highly successful Methods in Molecular Biology series, chapters include introductions to their respective topics, lists of the necessary materials and reagents, step-by-step, readily reproducible laboratory protocols, and tips on troubleshooting and avoiding known pitfalls. Authoritative and up-to-date, Molecular Plant Taxonomy: Methods and Protocols, Second Edition is an ideal guide for researchers seeking a better understanding of evolutionary processes, at species and population level, through molecular techniques.

The protein molecule is the basic building block of every living entity. Its deficiency leads to restricted growth and development of individuals. Globally, such malnutrition is on the rise due to various reasons such as rapid population growth, stagnation of productivity, and ever-rising costs. Millions of people, especially in developing and under-developed countries, suffer from protein malnutrition and the only possible solution is to encourage farmers to grow high-protein food legume crops in their fields for domestic consumption. This, however, could be possible if farmers are provided with new cultivars with high yield, and resistance to major insects, diseases, and key abiotic stresses. The major food legume crops are chickpea, cowpea, common bean, groundnut, lentil, pigeonpea, and soybean. Predominantly, the legume crops are grown under a subsistence level and, therefore, in comparison to cereals and horticultural crops their productivity is low and highly variable. The crop breeders around the globe are engaged in breeding suitable cultivars for harsh and changing environments but success has been limited and not up to needs. With the recent development of new technologies in plant sciences, efforts are being made to help under-privileged farmers through breeding new cultivars which will produce more protein per unit of land area. In this book, the contributors analyze the constraints, review new technologies, and propose a future course of crop breeding programs in seven cold and warm season legume crops.

Cryptic species are organisms which look identical, but which represent distinct evolutionary lineages. They are an emerging trend in organismal biology across all groups, from flatworms, insects, amphibians, primates, to vascular plants. This book critically evaluates the phenomenon of cryptic species and demonstrates how they can play a valuable role in improving our understanding of evolution, in particular of morphological stasis. It also explores how the recognition of cryptic species is intrinsically linked to the so-called 'species problem', the lack of a unifying species concept in biology, and suggests alternative approaches. Bringing together a range of perspectives from practicing taxonomists, the book presents case studies of cryptic species across a range of animal and plant groups. It will be an invaluable text for all biologists interested in species and their delimitation, definition, and purpose, including undergraduate and graduate students and researchers.

Phylogenetic reconstruction--the method by which biologists examine the relationship between living and extinct organisms in an effort to identify evolutionary pathways--has seen radical changes in the last ten years. But as rapid advances in mathematical, molecular, developmental, and cladistic techniques have greatly improved reconstruction efforts, they have also sparked controversy, especially with regard to the assumptions and data underlying the cladistic method. This book brings together contributions from a wide range of practitioners in order to find common ground. By examining numerous models from a variety of fields, the book serves as a guide to the latest methods of classification and phylogeny reconstruction and provides insights into the relative merits that each approach has to offer. In particular, developmental studies emerge as a significant factor in cladogram construction. Splendidly written and wide-ranging in scope, Models in Phylogeny Reconstruction will be welcomed by students and researchers in systematic and evolutionary biology.

To be published in 14 volumes over the next 12 years, the Flora of North America is an indispensable working resource for anyone with an interest in the distribution, habitat, morphology, and survival of the wide-ranging plant life in North America.

This introductory volume presents information on the physical and biological factors that have affected the evolution and distribution of plant life of North America. Current knowledge on geography, paleofloristics, and human impact is summarized, distilling the original field work of hundreds of contributors.

Beautifully illustrated, almost 100 drawings accompany the introductory essays, taken from various 19th and 20th century works. Chosen to illustrate plants mentioned within the text, they are also historical examples of the style and quality of botanical illustration from an earlier era to the present.

The complete collection will be an indispensable authoritative work for all botanical libraries and departments.

Starting with concise species accounts for all the marmoset and tamarin monkeys, this important new book then goes on to review their geographical distributions and still-contested taxonomy, along with comparative reviews of vocalizations, scent-marking, mating systems, infant care and development, social organization, and behaviour and ecology in the wild. As several of these small primates are rare or threatened, these subjects are strongly relevant to their management in captivity as well as for understanding natural populations. This is the first volume for several years to review current knowledge of this family, which comprises 52 species and subspecies found from Panama to northeastern Paraguay to southern Brazil.

Tropical lichens are not only of interest in themselves but are of importance in the study of the ecosystems of which they are a part. Information on tropical lichens has not hitherto been readily available in book form, and the object of this volume is to synthesize what is known at present and to indicate areas where further research is needed. The material presented here is derived from a meeting held by the Systematics Association at which contributions were presented on the systematic, ecological, and biogeographical aspects of tropical lichens, together with the problems of their conservation. The resulting volume should thus be of value to a wide variety of scientists.

A systematic treatise on the Chrysididae, a globally distributed family of wasps, also known as gold wasps (for their bright metallic colours) or cuckoo wasps (for their parasitic habits). Some 3,000 valid species have been named and are arranged in 84 genera and 4 sub-families. This book is the first re-evaluation of their taxonomy since 1889 and its four main goals are: to provide a worldwide overview of the family, with re-classification of the generic and higher taxa; to summarize previously published information; to indicate problems in need of further study; to give detailed synonymic species lists for each genus, where synonymy refers to scientific names used in different nomenclature systems to designate the same species. Discussions for each tribe and sub-family include ancestral characteristics, phylogenetically important characters and a corresponding cladogram, keys to genera, and relationships amongst taxa. Generic discussions include generic synonymy and diagnostic features, relationships to other genera, and detailed species lists. The book should be of interest to entomologists, taxonomists and systematists, especially in academic departments, natural history museum

This book examines sociobiology and evolution from the scientific and philosophical perspectives. Both sociobiology and evolution are areas of modern biology fraught with controversy and misunderstanding, yet fundamental to a coherent view of human life. For scientists and philosophers of science, at issue are the basic underpinnings of biology: explanation, determination, teleology, reductionism, and hierarchy. Professor Dyke describes the controversies involved, and argues that progress in sociobiology and evolution is hindered by an outmoded philosophical view of science - one that does not adequately take into account recent advances in our understanding of basic biological processes. The author aims to shift the focus from a philosophical understanding of biology to a dynamic, philosophically aware science.

This is the third of three volumes of "Keys to the Trematoda," a series on the systematics and identification of the Class Trematoda. It covers five superfamilies with the Order "Plagiorchiida" and the family "Didymozoidae," with the keys for their identification at the family, subfamily and generic levels. It also includes a key to all dignean superfamilies, including those treated in detail in volumes one and two.