In this Element, we extend our earlier treatment of biology's first
law. The law says that in any evolutionary system in which there is
variation and heredity, there is a tendency for diversity and
complexity to increase. The law plays the same role in biology that
Newton's first law plays in physics, explaining what biological
systems are expected to do when no forces act, in other words, what
happens when nothing happens. Here we offer a deeper explanation of
certain features of the law, develop a quantitative version of it,
and explore its consequences for our understanding of diversity and
complexity.
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