At first glance, studying behavior is easy, but as every budding ethologist quickly realizes, there are a host of complex practical, methodological and analytical problems to solve before designing and conducting the study. How do you choose which species or which behavior to study? What equipment will you need to observe and record behavior successfully? How do you record data in the dark, in the wet, or without missing part of the action? How do you analyze and interpret the data to yield meaningful information? This new expanded edition of the Handbook of Ethological Methods provides a complete step-by-step introduction to ethological methods from topic choice and behavioral description to data collection and statistical analysis. This book is a must for both beginning students and experienced researchers studying animal behavior in the field or laboratory.

This book is about the inner workings of one of nature’s most complex animal societies: the honey bee colony. It describes and illustrates the results of more than fifteen years of elegant experimental studies conducted by the author. In his investigations, Thomas Seeley has sought the answer to the question of how a colony of bees is organized to gather its resources. The results of his research—including studies of the shaking signal, tremble dance, and waggle dance, and other, more subtle means by which information is exchanged among bees—offer the clearest, most detailed picture available of how a highly integrated animal society works. By showing how several thousand bees function together as an integrated whole to collect the nectar, pollen, and water that sustain the life of the hive, Seeley sheds light on one of the central puzzles of biology: how units at one level of organization can work together to form a higher-level entity. In explaining why a hive is organized the way it is, Seeley draws on the literature of molecular biology, cell biology, animal and human sociology, economics, and operations research. He compares the honey bee colony to other functionally organized groups: multicellular organisms, colonies of marine invertebrates, and human societies. All highly cooperative groups share basic problems: of allocating their members among tasks so that more urgent needs are met before less urgent ones, and of coordinating individual actions into a coherent whole. By comparing such systems in different species, Seeley argues, we can deepen our understanding of the mechanisms that make close cooperation a reality.

With its iconic appearance and historic popular appeal, the giraffe is the world's tallest living terrestrial animal and the largest ruminant. Recent years have seen much-needed new research undertaken to improve our understanding of this unique animal. Drawing together the latest research into one resource, this is a detailed exploration of current knowledge on the biology, behaviour and conservation needs of the giraffe. Dagg highlights striking new data, covering topics such as species classification, the role of infrasound in communication, biological responses to external temperature changes and motherly behaviour and grief. The book discusses research into behaviour alongside practical information on captive giraffe, including diet, stereotypical behaviour, ailments and parasites, covering both problems and potential solutions associated with zoo giraffe. With giraffe becoming endangered species in Africa, the book ultimately focuses on efforts to halt population decline and the outlook for conservation measures.

With a new afterword, Why You Are Here: A speech on the opening of the COP26 climate summit As a young man, I felt I was out there in the wild, experiencing the untouched natural world - but it was an illusion. The tragedy of our time has been happening all around us, barely noticeable from day to day - the loss of our planet's wild places, its biodiversity. I have been witness to this decline. A Life on Our Planet contains my witness statement, and my vision for the future - the story of how we came to make this, our greatest mistake, and how, if we act now, we can yet put it right. We have the opportunity to create the perfect home for ourselves and restore the wonderful world we inherited. All we need is the will do so.

Additional resources for this book can be found at: www.wiley.com/go/davies/behaviouralecology

This textbook helped to define the field of Behavioural Ecology. In this fourth edition the text has been completely revised, with new chapters and many new illustrations and full colour photographs. The theme, once again, is the influence of natural selection on behaviour - an animal's struggle to survive and reproduce by exploiting and competing for resources, avoiding predators, selecting mates and caring for offspring, - and how animal societies reflect both cooperation and conflict among individuals.

Stuart A. West has joined as a co-author bringing his own perspectives and work on microbial systems into the book.Written in the same engaging and lucid style as the previous editions, the authors explain the latest theoretical ideas using examples from micro-organisms, invertebrates and vertebrates. There are boxed sections for some topics and marginal notes help guide the reader. The book is essential reading for students of behavioural ecology, animal behaviour and evolutionary biology.Key Features: Long-awaited new edition of a field-defining textbookNew chapters, illustrations and colour photographsNew co-authorFocuses on the influence of natural selection on behavior, and how animal societies reflect both cooperation and conflict among individuals

"The long-awaited update to a classic in this field is now here, presenting new direc-tions in thinking and addressing burning questions. Richly informed by progress in many other disciplines, such as sensory physiology, genetics and evolutionary theory, it marks the emergence of behav-ioural ecology as a fully fledged discipline..... This is a marvellous book, written in a lucid style. A must-read for those in the field, it is also a cornucopia of new thinking for anyone interested in evolution and behaviour."

"Manfred Milinski, "Nature, 2012

Speech, verbally mediated consciousness, and culture most evidently differentiate people from animals. Moreover, verbal communication has historically been viewed as a human attribute. Yet, the biological precursors to verbal communication exist in animal communication: obvious links have been found between human and animal nonverbal vocal communication. Knowledge of these links contribute to our understanding of three major theoretical and clinical issues: the evolution of vocal communication; the process of speech acquisition during infancy; and the pathways of disorders in communicative development. In Nonverbal Vocal Communication specialists from several disciplines review the present knowledge on neural substrates of vocal communication, on primate vocal communication, and on precursors and prerequisites of human speech. Among other points, the book illustrates that animal vocal signals, particularly in primates, appear to be much more complex than mere expressions of effective states. The nonverbal communication data is complemented by comparative and developmental research of the preverbal period of human vocal communication. Studies of speech acquisition, biological adaptation and environmental support to speech yield a complex picture of interrelations between biogenetic and environmental factors. The new evidence that parents--without knowing--are well-fitted for supporting infant communicative development is astonishing. Their capacities represent a primary, biological model of didactic educational support.

Carnivores have always fascinated us, even though they make up only 10% of all mammalian genera and only about 2% of all mammalian biomass. In Greek mythology most of the gods adorned their robes and helmets with depictions of carnivores, and the great hero Hercules' most famous feat was killing the "invulnerable" lion with his bare hands. Part* of our fascination with carnivores stems from fright and intrigue, and sometimes even hatred because of our direct competition with them. Cases of "man-eating" lions, bears, and wolves, as well as carnivores' reputation as killers of livestock and game, provoke communities and governrpents to adopt sweeping policies to exterminate them. Even President Theodore Roosevelt, proclaimer of a new wildlife protectionism, described the wolf as "the beast of waste and desolation. " The sheer presence and power of carnivores is daunt ing: they can move quickly yet silently through forests, attaining rapid bursts of speed when necessary; their massive muscles are aligned to deliver powerful attacks, their large canines and strong jaws rip open carcasses, and their scis sor-like carnassials slice meat. Partly because of our fear of these attributes, trophy hunting of carnivores has been, and to a certain extent still is, a sign of bravery and skill. Among some Alaskan Inuit, for example, a man is not eligible for marriage until he has killed a succession of animals of increasing size and dangerousness, culminating with the most menacing, the polar bear.

Through a global and interdisciplinary lens, this book discusses, analyzes and summarizes the novel conservation approach of rewilding. The volume introduces key rewilding definitions and initiatives, highlighting their similarities and differences. It reviews matches and mismatches between the current state of ecological knowledge and the stated aims of rewilding projects, and discusses the role of human action in rewilding initiatives. Collating current scholarship, the book also considers the merits and dangers of rewilding approaches, as well as the economic and socio-political realities of using rewilding as a conservation tool. Its interdisciplinary nature will appeal to a broad range of readers, from primary ecologists and conservation biologists to land managers, policy makers and conservation practitioners in NGOs and government departments. Written for a scientifically literate readership of academics, researchers, students, and managers, the book also acts as a key resource for advanced undergraduate and graduate courses.

Control theory can be roughly classified as deterministic or stochastic. Each of these can further be subdivided into game theory and optimal control theory. The central problem of control theory is the so called constrained maximization (which- with slight modifications--is equivalent to minimization). One can then say, heuristically, that the major problem of control theory is to find the maximum of some performance criterion (or criteria), given a set of constraints. The starting point is, of course, a mathematical representation of the performance criterion (or criteria)- sometimes called the objective functional--along with the constraints. When the objective functional is single valued (Le., when there is only one objective to be maximized), then one is dealing with optimal control theory. When more than one objective is involved, and the objectives are generally incompatible, then one is dealing with game theory. The first paper deals with stochastic optimal control, using the dynamic programming approach. The next two papers deal with deterministic optimal control, and the final two deal with applications of game theory to ecological problems. In his contribution, Dr. Marc Mangel applies the dynamic proQramming approach, as modified by his recent work--with Dr. Colin Clark, from the University of British Columbia (Mangel and Clark 1987}*--to modelling the "behavioral decisions" of insects. The objective functional is a measure of fitness. Readers interested in detailed development of the subject matter may consult Mangel (1985). My contributions deal with two applications of optimal control theory."

From the world-famous expert on chimpanzees - the powerfully compelling sequel to the international bestseller IN THE SHADOW OF MAN: 'An instant animal classic' Time Equipped with little more than a notebook, binoculars, and her fascination with wildlife, Jane braved a realm of unknowns to give the world a remarkable window into humankind's closest living relatives. On the shores of Lake Tanganyika, Gombe is a community where the principal residents are chimpanzees. Through Goodall's eyes we watch as the younger chimpanzees vie for power, and how the leaders must deal with this challenge. We learn how one mother successfully rears her children, whilst another appears to doom her offspring to failure. All life is here - glorious births and heart-breaking deaths, moments of brutality, alongside the most tender displays of affection. In THROUGH A WINDOW, as Jane Goodall reveals the story of this intimately intertwined community, we are shown the parallels with human emotions laid bare. Indeed, in the mirror of chimpanzee life, we see ourselves reflected.

The view of nature as `red in tooth and claw', as a jungle in which competition and predation are the predominant themes, has long been important in both the scientific and popular literature. However, in the past decade another view has become widespread among ecologists: the idea that mutualisms--mutually beneficial interactions between species--are just as important as competition and predation. This book is one of the first to explore this theme. Ideas and theories applicable to all sorts of mutualisms are presented and, where appropriate, examined in the light of concrete data. Themes explored include: the organisms involved, both animal and plant; how specializations evolved once mutualisms formed; how mutualisms affect population dynamics and community structure; and the role of mutualisms in different environments. The book will be of special interest to ecologists and a wide range of biologists.

Organized under broad ecological headings, this book provides a representative selection of present knowledge of mammal ecology with examples selected from a wide range of species and from all parts of the world.

Insect Conservation: A Global Synthesis is a landmark, field-defining work written by Professor Michael Samways, one of the founding fathers of this burgeoning discipline of conservation science. This book presents a state-of-the-art, comprehensive review of the entire field of insect conservation, from single-species conservation to whole-ecosystem approaches, and from natural ecosystems to the urban landscape. The variety and number of insects are truly vast. They are the most speciose group of organisms on Earth, with the majority barely known to science or still not described at all. They are a vital component in all terrestrial ecosystems, which would fail to function normally without them. Insect populations worldwide, however, are under threat. Human-mediated degradation of natural habitats, pollution, over-use of pesticides, and the spread of urbanization has led to the collapse of insect populations in many areas of the world. A growing recognition of the importance of insects in natural and agricultural systems has stimulated the development of an entire discipline dedicated to their conservation. Insect Conservation: A Global Synthesis is designed to be used by students of conservation biology and ecology, but also serves as an essential overview for professional entomologists with an interest in conservation, and for conservationists interested in insects. The book communicates on three levels: (i) through the text, with extensive references providing a gateway to the ever-increasing primary literature; (ii) through the extensive use of carefully constructed illustrations, with detailed captions which act to summarize the text and are complete in their own right; and (iii) through focused key points at the end of every chapter, which summarise the main learning points for students.

Wind farms are an essential component of global renewable energy policy and the action to limit the effects of climate change. There is, however, considerable concern over the impacts of wind farms on wildlife, leading to a wide range of research and monitoring studies, a growing body of literature and several international conferences on the topic. This unique multi-volume work provides a comprehensive overview of the interactions between wind farms and wildlife. Volume 1 documents the current knowledge of the potential impacts upon wildlife during both construction and operation. An introductory chapter on the nature of wind farms and the impact assessment process is followed by a series of in-depth chapters documenting effects on climatic conditions, vegetation, terrestrial invertebrates, aquatic invertebrates and fish, reptiles and amphibians, birds, bats and terrestrial mammals. A synopsis of the known and potential effects of wind farms upon wildlife in perspective concludes the volume. The authors have been carefully selected from across the globe from the large number of academics, consultants and practitioners now engaged in wind farm studies, for their influential contribution to the science. Edited by Martin Perrow and with contributions by 40 leading researchers including: Robert Barclay, Michael Dillon, Jan Olof Helldin, Hermann Hoetker, Jeffrey Lovich, Manuela de Lucas and Eugene Takle. The authors represent a wide range of organisations and institutions including the Universities of Calgary, Iowa State, Lund & Wyoming, US Geological Survey, Michael-Otto-Institut im NABU, Norwegian Institute for Nature Research, Spanish Council for Scientific Research, Renewable Energy Systems and several leading consultancies. Each chapter includes informative figures, tables, colour photographs and detailed case studies. Many of the latter are produced stand-alone from invited additional authors to ensure geographic spread and to showcase exciting new, often previously unpublished research. This book is designed for practitioners, researchers, managers and for a range of students in higher education, particularly those involved with environmental, ecological, conservation, impact assessment and climate change studies. Other volumes: Volume 2: Onshore: Monitoring and Mitigation (978-1-78427-123-7) Volume 3: Offshore: Potential Effects (978-1-78427-127-5) Volume 4: Offshore: Monitoring and Mitigation (978-1-78427-131-2)

A large number of comprehensive publications has been devoted to the Antarctic, to its plant and animal life. It is therefore relatively easy to familiarize oneself with the current state of Antarctic research. Nothing comparable is available for the Arctic. The heterogeneity and richness of the northern polar regions seem to have discouraged any attempt at a synthethic approach. This book has evolved from an attempt to summarize the results of 15 years of ecological and physiological research work in the Arctic - mostly on Spitsbergen. The necessity of comparing our results and the ecological conditions of Spitsbergen with other arctic regions grew into a full-sized book on arctic animal ecology. It is not meant as an exhaustive survey ofthe relevant literature. Instead I have tried to show how closely the various fields of research are interwoven, how many questions can be solved if only notice is taken of fellow scientists and their results, and how much arctic animals have in common. This book would not have been possible without the helpfulness of many colleagues. Above all I should like to mention Professor Ronning and Professor Solem of Trondheim University (Norway), Professor Arnthor Gardasson of Reykjavik University (Iceland), Dr. NettIeship, Dr. Oliver and Dr. Ryder of Canada and Professor West of Fairbanks University (Alaska, USA).

Symposium on Behaviour and Ecology, Saarbrucken, FRG, 1979"

n this book I have tried to bring together ideas from three different Istreams of thought with which I have been closely concerned during the last thirty years or so. The first is faunal history, usually regarded as a purely academic subject, but to some of whose events can be traced a number of the serious dislocations taking place in the world today. The second is ecology, particularly the structure and dynamics of populations. The third is conservation. I first published a few ideas about the signifi cance of invasions in 1943, in a war-time review called Polish Science and Learning, under the title of 'The changing realms of animal life'. Since then I have had the opportunity to think pretty hard about conservation, while taking part in the planning and development of the Nature Con servancy. In March 1957 I gave three broadcasts in the B. B. C. 's Third Programme, under the title of 'Balance and Barrier'. These were sub sequently printed in The Listener (1957, Vol. 57, pp. 514-15,556-7,596-7, and 600). The present book is essentially an expansion of these. I am extremely grateful to Mr James C. Thornton and Dr John Simons for in planning and giving these talks. advice and help In preparing this book I have had invaluable assistance from the staff of the Bureau of Animal Population. Miss C. M. Gibbs typed the fair copy. Miss M."

The study of animal development has deep historical roots in codifying the field of evolutionary biology. In the 1940s, evolutionary theory became engulfed by microevolutionary genetic analysis and development became focused on mechanisms, forsaking the evolutionary implications of ontogeny. Recently, ontogeny has resurfaced as a significant component of evolutionary change, population, and community dynamics. Ontogeny, Functional Ecology and Evolution of Bats is a unique reference work by bat biologists who emphasize the importance of understanding ontogeny in the analysis of evolution and ecology. In addition, the developmental underpinnings of specialized morphology, physiology, and behavior are elucidated, and the strong influence of ecology on the ontological niche of juvenile bats is illustrated. This book is an essential reference for bat biologists, and all those working in the fields of ecology, developmental biology, evolution, behavior, and systematics.

A witty and informative account that busts the myths about Britain's most captivating butterfly species. When summer is at its zenith and the sallow foliage develops a bluish tinge, a giant butterfly – beautiful, bold and brazen – flies powerfully over the tree canopy. Females of this species, wary yet determined, haunt the sallow thickets, depositing their eggs, while the males establish treetop territories and descend to the woodland floor in search of indelicacies to feed upon. Mysterious, elusive and enthralling in equal measure, this is the butterfly that Victorian collectors yearned for above all others: His Imperial Majesty, the Purple Emperor. A wondrous enigma, the Purple Emperor is our most elusive and least-known butterfly – we glimpse it only through fissures in its treetop world, yet this giant insect has fascinated us for centuries and has even inspired its own 'Emperoring' language. Matthew Oates became captivated by the Purple Emperor following his first sighting as a boy. He has studied it assiduously ever since, devoting his life to trying to unravel the Emperor's secrets. His Imperial Majesty takes us on a journey, beginning with a dalliance into the bizarre history of our engagement with the butterfly, with daring doings and gross eccentricities from the Victorian and Edwardian eras. Subsequent chapters explore all aspects of this remarkable butterfly's life cycle, including behaviour, habitat preferences, life history and conservation, all relayed in Matthew's unique, informative and witty style. Not so long ago, our knowledge of the Emperor was largely based on a blend of mythology and assumption. This book dispels the fabrications and reveals all about the Purple Emperor – the king of British butterflies.

Winner of the Richard Jefferies Society and White Horse Book Shop Literary Prize 'splendid' -Guardian 'visionary' -New Statesman Britain has all the space it needs for an epic return of its wildlife. Only six percent of our country is built upon. Contrary to popular myth, large areas of our countryside are not productively farmed but remain deserts of opportunity for both wildlife and jobs. It is time to turn things around. Praised as 'visionary' by conservationists and landowners alike, Rebirding sets out a compelling manifesto for restoring Britain's wildlife, rewilding its species and restoring rural jobs - to the benefit of all.

The past decade has seen a huge increase in the interest and attention directed toward sea ducks, the Mergini tribe. This has been inspired, in large part, by the conservation concerns associated with numerical declines in several sea duck species and populations, as well as a growing appreciation for their interesting ecological attributes. Reflecting the considerable research recently conducted on this tribe, Ecology and Conservation of North American Sea Ducks examines the 15 extant species of sea ducks from North America. Chapters are organized conceptually to focus on, compare, and contrast the ecological attributes of the tribe. Experts provide in-depth treatments of a range of topics, including: Population dynamics and genetics Infectious diseases and parasites Breeding costs and cross-seasonal affects Contaminant burdens Foraging behavior and energetics Migration strategies, molt ecology, and habitat affinities and dynamics Breeding systems and reproductive behavior Harvest history The book presents a comprehensive synthesis of sea duck ecology, documents factors that have caused population declines of some species, and provides managers with measures to enhance recovery of depressed populations of sea ducks in North America. Capturing the current state of knowledge of this unique tribe, it provides a benchmark for where we are in conservation efforts and suggests future directions for researchers, managers, students, conservationists, and avian enthusiasts.

The most up-to-date and authoritative resource on the biology and evolution of solitary bees While social bees such as honey bees and bumble bees are familiar to most people, they comprise less than 10 percent of all bee species in the world. The vast majority of bees lead solitary lives, surviving without the help of a hive and using their own resources to fend off danger and protect their offspring. This book draws on new research to provide a comprehensive and authoritative overview of solitary bee biology, offering an unparalleled look at these remarkable insects. The Solitary Bees uses a modern phylogenetic framework to shed new light on the life histories and evolution of solitary bees. It explains the foraging behavior of solitary bees, their development, and competitive mating tactics. The book describes how they construct complex nests using an amazing variety of substrates and materials, and how solitary bees have co-opted beneficial mites, nematodes, and fungi to provide safe environments for their brood. It looks at how they have evolved intimate partnerships with flowering plants and examines their associations with predators, parasites, microbes, and other bees. This up-to-date synthesis of solitary bee biology is an essential resource for students and researchers, one that paves the way for future scholarship on the subject. Beautifully illustrated throughout, The Solitary Bees also documents the critical role solitary bees play as crop pollinators, and raises awareness of the dire threats they face, from habitat loss and climate change to pesticides, pathogens, parasites, and invasive species.

This book is about the ways in which many animals form groups; for instance, schools of fish, flocks of birds, and swarms of insects. Covering both invertebrate and vertebrate species, the authors investigate three-dimensional animal aggregations from a variety of disciplines, from physics to mathematics to biology. The first section is devoted to the various methods, mainly optical and acoustic, used to collect three-dimensional data over time. The second section focuses on analytical methods used to quantify pattern, group kinetics, and interindividual interactions within the group. The section on behavioural ecology and evolution deals with the functions of aggregative behaviour from the point of view of an inherently selfish individual member. The final section uses models to elucidate how group dynamics at the individual level creates emergent pattern at the level of the group.

These charismatic mammals, which include dogs, cats, hyenas, weasels, mongooses, seals, sea lions and bears, have always held special importance to humans throughout history and continue to do so today. In recent decades, the emergence of new technologies has completely transformed our knowledge of how carnivorans interact with their environments and consequently reshaped our view of carnivoran ecology. This unique synthesis uses examples from a diverse and expanding carnivoran literature, drawing from all carnivoran families and spanning the world's oceans and continents, to produce a clearly written and richly illustrated book that reviews our current state of knowledge of carnivoran ecology. It addresses all levels of biological organization and function, from genes to enzymes, organisms, populations, and ecosystems. Special attention is given to how carnivoran species interact with their prey, each other, and humans. There is an emphasis on community interactions and their importance in carnivoran evolution, showing how evolutionary constraints (morphological, physiological, and behavioral) structure communities today. The book's approach is strongly comparative, contrasting herbivores with carnivores, predators with scavengers, and cats with dogs. Carnivorans play important roles in many high-profile conservation cases, either as species of concern or agents of endangerment, and their importance is demonstrated in both contexts. Carnivoran Ecology is an accessible advanced textbook aimed principally at senior undergraduate and graduate students taking courses in carnivore ecology, as well as a broad audience of professional academics (especially carnivore and mammalian biologists), researchers, and practitioners working in both governmental and non-governmental organizations. A significant secondary market will exist amongst the large amateur naturalist community including those wishing to explore the ecological and evolutionary links between domestic carnivorans (dogs, cats, ferrets etc.) and their wild counterparts.

Visual ecology is the study of how animals use visual systems to meet their ecological needs, how these systems have evolved, and how they are specialized for particular visual tasks. "Visual Ecology" provides the first up-to-date synthesis of the field to appear in more than three decades. Featuring some 225 illustrations, including more than 140 in color, spread throughout the text, this comprehensive and accessible book begins by discussing the basic properties of light and the optical environment. It then looks at how photoreceptors intercept light and convert it to usable biological signals, how the pigments and cells of vision vary among animals, and how the properties of these components affect a given receptor's sensitivity to light. The book goes on to examine how eyes and photoreceptors become specialized for an array of visual tasks, such as navigation, evading prey, mate choice, and communication.

A timely and much-needed resource for students and researchers alike, "Visual Ecology" also includes a glossary and a wealth of examples drawn from the full diversity of visual systems.The most up-to-date overview of visual ecology availableFeatures some 225 illustrations, including more than 140 in color, spread throughout the textGuides readers from the basic physics of light to the role of visual systems in animal behaviorIncludes a glossary and a wealth of real-world examples