Living things are organized in a hierarchy of levels. Genes group together in cells, cells group together in organisms, and organisms group together in societies. Even different species form mutualistic partnerships. Throughout the history of life, previously independent units have formed groups that, in time, have come to resemble individuals in their own right. Evolutionary biologists term such events "the major transitions." The process common to them all is social evolution. Each transition occurs only if natural selection favors one unit joining with another in a new kind of group.
This book presents a fresh synthesis of the principles of social evolution that underlie the major transitions, explaining how the basic theory underpinning social evolution - inclusive fitness theory - is central to understanding each event. The book defines the key stages in a major transition, then highlights the shared principles operating at each stage across the transitions as a whole. It addresses in new ways the question of how, once they have arisen, organisms and societies become more individualistic.

Living things are organized in a hierarchy of levels. Genes group together in cells, cells group together in organisms, and organisms group together in societies. Even different species form mutualistic partnerships. Throughout the history of life, previously independent units have formed groups that, in time, have come to resemble individuals in their own right. Evolutionary biologists term such events "the major transitions." The process common to them all is social evolution. Each transition occurs only if natural selection favors one unit joining with another in a new kind of group.
This book presents a fresh synthesis of the principles of social evolution that underlie the major transitions, explaining how the basic theory underpinning social evolution - inclusive fitness theory - is central to understanding each event. The book defines the key stages in a major transition, then highlights the shared principles operating at each stage across the transitions as a whole. It addresses in new ways the question of how, once they have arisen, organisms and societies become more individualistic.

Additional resources for this book can be found at: www.wiley.com/go/davies/behaviouralecology

This textbook helped to define the field of Behavioural Ecology. In this fourth edition the text has been completely revised, with new chapters and many new illustrations and full colour photographs. The theme, once again, is the influence of natural selection on behaviour - an animal's struggle to survive and reproduce by exploiting and competing for resources, avoiding predators, selecting mates and caring for offspring, - and how animal societies reflect both cooperation and conflict among individuals.

Stuart A. West has joined as a co-author bringing his own perspectives and work on microbial systems into the book.Written in the same engaging and lucid style as the previous editions, the authors explain the latest theoretical ideas using examples from micro-organisms, invertebrates and vertebrates. There are boxed sections for some topics and marginal notes help guide the reader. The book is essential reading for students of behavioural ecology, animal behaviour and evolutionary biology.Key Features: Long-awaited new edition of a field-defining textbookNew chapters, illustrations and colour photographsNew co-authorFocuses on the influence of natural selection on behavior, and how animal societies reflect both cooperation and conflict among individuals

"The long-awaited update to a classic in this field is now here, presenting new direc-tions in thinking and addressing burning questions. Richly informed by progress in many other disciplines, such as sensory physiology, genetics and evolutionary theory, it marks the emergence of behav-ioural ecology as a fully fledged discipline..... This is a marvellous book, written in a lucid style. A must-read for those in the field, it is also a cornucopia of new thinking for anyone interested in evolution and behaviour."

"Manfred Milinski, "Nature, 2012

Introducing Biological Energetics is a novel, interdisciplinary text that presents biological understanding in terms of general underlying principles, treating energy as the overarching theme and emphasizing the all-pervading influence of energy transformation in every process, both living and non-living. Key processes and concepts are explained in turn, culminating in a description of the overall functioning and regulation of a living cell. The book rounds off the story of life with a brief account of the endosymbiotic origins of eukaryotic cells, the development of multicellularity, and the emergence of modern plants and animals.
Multidisciplinary research in science is becoming commonplace. However, as traditional boundaries start to break down, researchers are increasingly aware of the deficiencies in their knowledge of related disciplines. Introducing Biological Energetics redresses the reciprocal imbalance in the knowledge levels of physical and biological scientists in particular. Its style of presentation and depth of treatment has been carefully designed to unite these two readerships.

Recommended in CHOICE, February 2021 Insects are all around us, outweighing humanity by 17 times. Many are nuisances; they compete with us for food and carry some of our most devastating diseases. Many common pests have been transported worldwide by humans. Yet, some recent reports suggest dramatic declines in some important groups, such as pollinators and detritivores. Should we care? Yes, we should. Without insect pollinators we'd lose 35% of our global food production; without detritivores, we would be buried in un-decayed refuse. Insects are also critical sources for nutritional, medical and industrial products. A world without insects would seem a very different and unpleasant place. So why do insects inspire such fear and loathing? This concise, full-color text challenges many entrenched perceptions about insect effects on our lives. Beginning with a summary of insect biology and ecology that affect their interactions with other organisms, it goes on to describe the various positive and negative ways in which insects and humans interact. The final chapters describe factors that affect insect abundance and approaches to managing insects that balance their impacts. The first textbook to cater directly to those studying Insect and Society or Insect Ecology modules, this book will also be fascinating reading for anyone interested in learning how insects affect human affairs and in applying more sustainable approaches to "managing" insects. This includes K-12 teachers, undergraduate students, amateur entomologists, conservation practitioners, environmentalists, as well as natural resource managers, land use planners and environmental policy makers.

Bioinvasions and Globalization synthesises our current knowledge of the ecology and economics of biological invasions, providing an in-depth evaluation of the science and its implications for managing the causes and consequences of one of the most pressing environmental issues facing humanity today.
Emergent zoonotic diseases such as HIV and SARS have already imposed major costs in terms of human health, whilst plant and animal pathogens have had similar effects on agriculture, forestry, fisheries. The introduction of pests, predators and competitors into many ecosystems has disrupted the benefits they provide to people, in many cases leading to the extirpation or even extinction of native species. This timely book analyzes the main drivers of bioinvasions - the growth of world trade, global transport and travel, habitat conversion and land use intensification, and climate change - and their consequences for ecosystem functioning. It shows how bioinvasions impose disproportionately high costs on countries where a large proportion of people depend heavily on the exploitation of natural resources. It considers the options for improving assessment and management of invasive species risks, and especially for achieving the international cooperation needed to address bioinvasions as a negative externality of international trade.

Community ecology is the study of the interactions between populations of co-existing species. Co-edited by two prominent community ecologists and featuring contributions from top researchers in the field, this book provides a survey of the state-of-the-art in both the theory and applications of the discipline. It pays special attention to topology, dynamics, and the importance of spatial and temporal scale while also looking at applications to emerging problems in human-dominated ecosystems (including the restoration and reconstruction of viable communities).
Community Ecology: Processes, Models, and Applications adopts a mainly theoretical approach and focuses on the use of network-based theory, which remains little explored in standard community ecology textbooks. The book includes discussion of the effects of biotic invasions on natural communities; the linking of ecological network structure to empirically measured community properties and dynamics; the effects of evolution on community patterns and processes; and the integration of fundamental interactions into ecological networks. A final chapter indicates future research directions for the discipline.

Bumblebees are familiar and charismatic insects, occurring throughout much of the world. They are increasingly being used as a model organism for studying a wide range of ecological and behavioural concepts, such as social organization, optimal foraging theories, host-parasite interactions, and pollination. Recently they have become a focus for conservationists due to mounting evidence of range contractions and catastrophic extinctions with some species disappearing from entire continents (e.g. in North America). Only by improving our understanding of their ecology can we devise sensible plans to conserve them. The role of bumblebees as invasive species (e.g. Bombus terrestris in Japan) has also become topical with the growing trade in commercial bumblebee nests for tomato pollination leading to establishment of non-native bumblebees in a number of countries.
Since the publication of the first edition of the book, there have been hundreds of research papers published on bumblebees. There is clearly a continuing need for an affordable, well-illustrated, and appealing text that makes accessible all of the major advances in understanding of the behaviour and ecology of bumblebees that have been made in the last 30 years.

Coastal zones are becoming increasingly topical (and politically sensitive) as they face relentless pressures from urban expansion, recreational development, and sea level rise due to climate change. This timely book provides a comprehensive introduction to the formation, dynamics, maintenance, and perpetuation of coastal sand dune systems. It describes the interactions between living organisms and the physical processes of geomorphology. A global range of examples enhance the book's international appeal. Based on the research presented in this book, simple to complex field studies and experiments could be designed at undergraduate and graduate levels to illustrate various biological principles. This accessible book is intended for a diverse audience; as an invaluable reference for researchers who study coastal dune systems and for novice researchers requiring a sound introduction to the subject. This book is suitable for both senior undergraduate and graduate students taking courses in coastal zone management, plant ecology, restoration ecology, and conservation biology, as well as the many professional ecologists and conservation biologists requiring a concise but authoritative overview of the topic. The book also will be of relevance and use to coastal managers, planners, naturalists, and anyone pursuing a greater understanding of coastal sand dunes.

Coastal zones are becoming increasingly topical (and politically sensitive) as they face relentless pressures from urban expansion, recreational development, and sea level rise due to climate change. This timely book provides a comprehensive introduction to the formation, dynamics, maintenance, and perpetuation of coastal sand dune systems. It describes the interactions between living organisms and the physical processes of geomorphology. A global range of examples enhance the book's international appeal. Based on the research presented in this book, simple to complex field studies and experiments could be designed at undergraduate and graduate levels to illustrate various biological principles.
This accessible book is intended for a diverse audience; as an invaluable reference for researchers who study coastal dune systems and for novice researchers requiring a sound introduction to the subject. This book is suitable for both senior undergraduate and graduate students taking courses in coastal zone management, plant ecology, restoration ecology, and conservation biology, as well as the many professional ecologists and conservation biologists requiring a concise but authoritative overview of the topic. The book also will be of relevance and use to coastal managers, planners, naturalists, and anyone pursuing a greater understanding of coastal sand dunes.

With the exception of climate change, biological invasions have probably received more attention during the past ten years than any other ecological topic. Yet this is the first synthetic, single-authored overview of the field since Williamson's 1996 book. Written fifty years after the publication of Elton's pioneering monograph on the subject, Invasion Biology provides a comprehensive and up-to-date review of the science of biological invasions while also offering new insights and perspectives relating to the processes of introduction, establishment, and spread. The book connects science with application by describing the health, economic, and ecological impacts of invasive species as well as the variety of management strategies developed to mitigate harmful impacts. The author critically evaluates the approaches, findings, and controversies that have characterized invasion biology in recent years, and suggests a variety of future research directions. Carefully balanced to avoid distinct taxonomic, ecosystem, and geographic (both investigator and species) biases, the book addresses a wide range of invasive species (including protists, invertebrates, vertebrates, fungi, and plants) which have been studied in marine, freshwater, and terrestrial environments throughout the world by investigators equally diverse in their origins.
This accessible and thought-provoking text will be of particular interest to graduate level students and established researchers in the fields of invasion biology, community ecology, conservation biology, and restoration ecology. It will also be of value and use to land managers, policy makers, and other professionals charged with controlling the negative impacts associated with recently arrived species.

Water is fundamental to life and all metabolic reactions are influenced by the aqueous environment in which they occur. Osmoregulation and water balance are therefore absolutely essential topics in animal physiology.
Animal Osmoregulation collates a widely dispersed literature to produce a comprehensive and authoritative synthesis of the field, providing detailed examples of osmoregulatory processes at the organismal, organ and cellular level. It incorporates clear background information on ion regulation and transport (specifically in the light of recent molecular studies) and illustrates the physical principles to which each organism must adhere, as well as the phylogenetic constraints within which it must operate.
As with other titles in the Oxford Animal Biology Series, the topic is addressed using examples from throughout the animal kingdom, identifying common themes that transcend taxonomy.

Wind farms are an essential component of global renewable energy policy and the action to limit the effects of climate change. There is, however, considerable concern over the impacts of wind farms on wildlife, leading to a wide range of research and monitoring studies, a growing body of literature and several international conferences on the topic. This unique multi-volume work provides a comprehensive overview of the interactions between wind farms and wildlife. Volume 3 documents the current knowledge of the potential effects upon wildlife during both construction and operation of offshore wind farms. An introductory chapter on the nature of wind farms and the legislation surrounding them is followed by a series of in-depth chapters documenting effects on physical processes, atmosphere and ocean dynamics, seabed communities, fish, marine mammals, migratory birds and bats and seabirds. A synopsis of the known and potential effects of wind farms upon wildlife concludes the volume. The authors have been carefully selected from across the globe from the large number of academics, consultants and practitioners now engaged in wind farm studies, for their influential contribution to the science. Edited by Martin Perrow and with contributions by 30 leading researchers including: Goeran Brostroem, Steven Degraer, Mike Elliot, Andrew Gill, Ommo Huppop, Georg Nehls and Nicolas Vanermen. The authors represent a wide range of organisations and institutions including the Universities of Gothenburg, Hamburg and Hull, Alfred Wegener Institute, Cefas (UK), Research Institute for Nature and Forest (INBO), Royal Belgian Institute of Natural Sciences, Vattenfall and several leading consultancies. Each chapter includes informative figures, tables, colour photographs and detailed case studies, including some from invited authors to showcase exciting new research. Other volumes: Volume 1: Onshore: Potential Effects (978-1-78427-119-0) Volume 2: Onshore: Monitoring and Mitigation (978-1-78427-123-7) Volume 4: Offshore: Monitoring and Mitigation (978-1-78427-131-2)

Understanding of animal social and sexual evolution has seen a renaissance in recent years with discoveries of frequent infidelity in apparently monogamous species, the importance of sperm competition, active female mate choice, and eusocial behavior in animals outside the traditional social insect groups. Each of these findings has raised new questions, and suggested new answers, about the evolution of behavioral interactions among animals. This volume synthesizes recent research on the sexual and social biology of the Crustacea, one of the dominant invertebrate groups on earth. Its staggering diversity includes ecologically important inhabitants of nearly every environment from deep-sea trenches, through headwater streams, to desert soils. The wide range of crustacean phenotypes and environments is accompanied by a comparable diversity of behavioral and social systems, including the elaborate courtship and wildly exaggerated morphologies of fiddler crabs, the mysterious queuing behavior of migrating spiny lobsters, and even eusociality in coral-reef shrimps. This diversity makes crustaceans particularly valuable for exploring the comparative evolution of sexual and social systems. Despite exciting recent advances, however, general recognition of the value of Crustacea as models has lagged behind that of the better studied insects and vertebrates. This book synthesizes the state of the field in crustacean behavior and sociobiology and places it in a conceptually based, comparative framework that will be valuable to active researchers and students in animal behavior, ecology, and evolutionary biology. It brings together a group of internationally recognized and rising experts in fields related to crustacean behavioral ecology, ranging from physiology and functional morphology, through mating and social behavior, to ecology and phylogeny. Each chapter makes connections to other, non-crustacean taxa, and the volume closes with a summary section that synthesizes the contributions, discusses anthropogenic impacts, highlights unanswered questions, and provides a vision for profitable future research.

Archie Carr, one of the greatest biologists of the twentieth century, played a leading part in finding a new and critical role for natural history and systematics in a post-1950s world dominated by the glamorous science of molecular biology. With the rise of molecular biology came a growing popular awareness of species extinction. Carr championed endangered sea turtles, and his work reflects major shifts in the study of ecology and evolution. A gifted nature writer, his books on the natural history of sea turtles and their habitats in Florida, the Caribbean, and Africa entertained and educated a wide audience. Carr's conservation ethic grew from his field work as well as his friendships with the fishermen who supplied him with many of the stories he retold so engagingly. With Archie Carr as the focus, The Man Who Saved Sea Turtles explores the evolution of the naturalist tradition, biology, and conservation during the twentieth century.

Human-wildlife conflict (HWC) is one of the most complex and urgent issues facing wildlife management and conservation today. Originally focused on the ecology and economics of wildlife damage, the study and mitigation of HWC has gradually expanded its scope to incorporate the human dimensions of the whole spectrum of human-wildlife relationships, from conflict to coexistence. Having the conflict-to-coexistence continuum as its leitmotiv, this book explores a variety of theories and methods currently used to address human-wildlife interactions, illustrated by case studies from around the world. It presents some key concepts in the field, such as values, emotions, social identity and tolerance, and a variety of insights and solutions to turn conflict into coexistence, from individual level to national scales, including conservation marketing, incremental and radical innovation, strategic planning, and socio-ecological systems. This volume will be of interest to a wide range of readers, including academics, researchers, students, practitioners and policy-makers.

The last documented sighting of the Ivory-billed Woodpecker--one of the rarest and most intriguing animals in the world--was noted over 50 years ago. Long thought to be extinct, the 2005 announcement of a sighting in Arkansas sparked tremendous enthusiasm and hope that this species could yet be saved. But the subsequent failure of a massive search to relocate Ivorybills in Arkansas made hope for the species' revival short-lived.
Here, noted ornithologist Geoffrey Hill tells the story of how he and two of his colleagues stumbled upon what may be a breeding population of Ivory-billed Woodpeckers in the swamps of northern Florida. He relates their laborious attempts to document irrefutable evidence for the existence of this shy, elusive bird following the failure of a much larger research team to definitively prove the bird's existence.
Hill tells of his travails both in and out of the vast swamp wilderness, pulling back the curtain to reveal the little-seen political maneuvering that is part of all modern science. He explains how he and his group decided who to exclude or include as their findings came in, and why they felt the need to keep their search a secret. Hill returns repeatedly to how expectations can guide observations, and how tempting it is to oversell evidence in the face of the struggle between an overwhelming desire to find the bird and the need to retain integrity and objectivity.
Written like a good detective story, Ivorybill Hunters also delves into the science behind the rediscovery of a species, explaining how professional ornithologists follow up on a sight record of a rare bird, and how this differs from the public's perception of how scientistsactually work. Hill notes the growing role of amateurs in documenting bird activity and discusses how the community of birders and nature lovers can see, enjoy, and help preserve these birds.
Ivorybill Hunters will prove a fascinating read for those with an interest in natural history, adventure, environmental conservation, and science, as well as the more than forty-six million Americans who now call themselves birdwatchers.

Why do baby sharks, hyenas, and pelicans kill their siblings? Why do beetles and mice commit infanticide? Why are twins and birth defects more common in older human mothers? "A Natural History of Families" concisely examines what behavioral ecologists have discovered about family dynamics and what these insights might tell us about human biology and behavior. Scott Forbes's engaging account describes an uneasy union among family members in which rivalry for resources often has dramatic and even fatal consequences.

In nature, parents invest resources and control the allocation of resources among their offspring to perpetuate their genetic lineage. Those families sometimes function as cooperative units, the nepotistic and loving havens we choose to identify with. In the natural world, however, dysfunctional familial behavior is disarmingly commonplace.

While explaining why infanticide, fratricide, and other seemingly antisocial behaviors are necessary, Forbes also uncovers several surprising applications to humans. Here the conflict begins in the moments following conception as embryos struggle to wrest control of pregnancy from the mother, and to wring more nourishment from her than she can spare, thus triggering morning sickness, diabetes, and high blood pressure. Mothers, in return, often spontaneously abort embryos with severe genetic defects, allowing for prenatal quality control of offspring.

Using a broad sweep of entertaining examples culled from the world of animals and humans, "A Natural History of Families" is a lively introduction to the behavioral ecology of the family.

This volume provides a modern introduction to the soil fauna and their contributions to ecosystem function, the mechanisms that structure soil fauna assemblages from local to global scales, and the potential impacts of global change on soil fauna assemblages and through this ecosystem function. Wanting to be an accessible primer, this book is a high level overview of current knowledge rather than a detailed tome of all existing information, with emphasis being placed on key findings and general patterns. It focuses on the soil fauna but contextualizes these assemblages in relation to the microbial assemblages belowground and the vegetation aboveground. It is clear that our knowledge of soil fauna assemblages is ever increasing, but there is still a lot to discover. Key areas of research are highlighted, with particular reference to the future of soil fauna assemblages.

Many infectious diseases of recent concern, including malaria, cholera, plague, and Lyme disease, have emerged from complex ecological communities, involving multiple hosts and their associated parasites. Several of these diseases appear to be influenced by human impacts on the environment, such as intensive agriculture, clear-cut forestry, and habitat loss and fragmentation; such environmental impacts may affect many species that occur at trophic levels below or above the hostcommunity. These observations suggest that the prevalence of both human and wildlife diseases may be altered in unanticipated ways by changes in the structure and composition of ecological communities. Predicting the epidemiological ramifications of such alteration in community composition will require strengthening the current union between community ecology and epidemiology.

Wildfowl and screamers belong to a highly diverse family of birds, confined to watery habitats. They are amongst the most attractive of birds and are very well-known to man, who has domesticated them, used their feathers for warm clothing and ornamentation, admired their flight, courtship and migration, caught them for food, maintained them in captivity for pleasure, and written about their doings in delightful children's stories, from Mother Goose to Jemima Puddleduck and Donald Duck. They occur throughout the world except Antarctica. Some are faithful to the same partner for life, others for only the few minutes of copulation. In some species, male and female make devoted parents, and yet there is one within the group whose female lays her eggs in the nests of others and never incubates. Diving as a method of obtaining food has evolved many times within the family. Most nest in the open but others in the tree-hole nests of woodpeckers and some in the ground burrows of rabbits or aardvarks. They may be highly social or solitary, defending a large territory.
Ducks, Geese, and Swans begins with eight chapters giving an overview of the family, their taxonomy and evolution, feeding ecology, breeding strategies, social behavior, movements and migrations, population dynamics, and conservation and management, followed by accounts of 165 species, written by a team of expert wildfowl specialists, describing each bird in its natural state and summarizing the published literature and recent research. Complementing the accounts are thirty specially commissioned color plates by Mark Hulme, along with numerous black and white drawings illustrating behaviors, plus distribution maps for each species.

From foraging patterns in a single tree to social interactions across a home range, how primates use space is a key question in the field of primate behavioral ecology. Drawing on the latest advances in spatial analysis tools, this book offers practical guidance on applying geographic information systems (GIS) to central questions in primatology. An initial methodological section discusses niche modelling, home range analysis and agent-based modelling, with a focus on remote data collection. Research-based chapters demonstrate how ecologists apply this technology to a suite of topics including: calculating the intensity of use of both range and travel routes, assessing the impacts of logging, mining and hunting, and informing conservation strategies.

Current dominant thinking and practice in the private and public sectors asserts that peoples' development needs are in conflict with, or mutually exclusive to, the need to conserve the biosphere on which we depend. Consequently, we are asked to either diminish development in the name of conservation or diminish conservation in the name of development. Efforts to identify complementary objectives, or mutually acceptable trade-offs and compromises indicate, however, that this does not always have to be the case. This first volume in the State of the Apes series draws attention to the evolving context within which great ape and gibbon habitats are increasingly interfacing with extractive industries. Intended for a broad range of policy makers, industry experts, decision makers, academics, researchers and NGOs, these publications aim to influence debate, practice and policy, seeking to reconcile ape conservation and welfare, and economic and social development, through objective and rigorous analysis.

With its iconic appearance and historic popular appeal, the giraffe is the world's tallest living terrestrial animal and the largest ruminant. Recent years have seen much-needed new research undertaken to improve our understanding of this unique animal. Drawing together the latest research into one resource, this is a detailed exploration of current knowledge on the biology, behaviour and conservation needs of the giraffe. Dagg highlights striking new data, covering topics such as species classification, the role of infrasound in communication, biological responses to external temperature changes and motherly behaviour and grief. The book discusses research into behaviour alongside practical information on captive giraffe, including diet, stereotypical behaviour, ailments and parasites, covering both problems and potential solutions associated with zoo giraffe. With giraffe becoming endangered species in Africa, the book ultimately focuses on efforts to halt population decline and the outlook for conservation measures.

The book contains a complete description of the natural history and biology of the Ratites and Tinamous - the group of flightless birds that includes ostriches, emus, cassowaries and kiwis. Each species is treated in turn, with seven general introductory chapters and full colour illustrations of all 55 species, as well as over 50 line drawings. The author has spent 25 years research on the ratites and has drawn together the extensive and often inaccessible literature on the groups.