At first glance, studying behavior is easy, but as every budding ethologist quickly realizes, there are a host of complex practical, methodological and analytical problems to solve before designing and conducting the study. How do you choose which species or which behavior to study? What equipment will you need to observe and record behavior successfully? How do you record data in the dark, in the wet, or without missing part of the action? How do you analyze and interpret the data to yield meaningful information? This new expanded edition of the Handbook of Ethological Methods provides a complete step-by-step introduction to ethological methods from topic choice and behavioral description to data collection and statistical analysis. This book is a must for both beginning students and experienced researchers studying animal behavior in the field or laboratory.

The Great Apes (chimpanzees, bonobos, gorillas and orangutans) are our closest living relatives, sharing a common ancestor only five million years ago. We also share key features such as high intelligence, omnivorous diets, prolonged child-rearing and rich social lives. The Great Apes show a surprising diversity of adaptations, particularly in social life, ranging from the solitary life of orangutans, through patriarchy in gorillas to complex but different social organizations in bonobos and chimpanzees. As Great Apes are so close to humans, comparisons yield essential knowledge for modeling human evolutionary origins. Great Ape Societies provides comprehensive up-to-date syntheses of work on all four species, drawing on decades of international field work, zoo and laboratory studies. It will be essential reading for students and researchers in primatology, anthropology, psychology and human evolution.

Alan Powderham's dazzling photography is complemented by Dr Sancia van der Meij's scientific expertise, creating a scientifically rigorous yet breathtakingly beautiful reference work. Van der Meij is an experienced coral reef specialist while Powderham's use of rebreather diving technology allows him to get unusually close to his subject matter. The spectacular landscapes and natural riches of the Coral Triangle are under threat from overfishing, climate change, unsustainable tourism, habitat destruction and poor governance. This book showcases exactly why we need to protect this unique ecosystem, contrasting its existing beauty with images of the damage already being perpetrated. 10% of the author royalties are going to Conservation International to aid their work. The book covers many of the groups of animals seen on a reef that have been largely omitted by others. It reports engaging observations giving insights to many unique animal behaviours and relationships, that make it stand out from other books.

Statistical Methods for Field and Laboratory Studies in Behavioral Ecology focuses on how statistical methods may be used to make sense of behavioral ecology and other data. It presents fundamental concepts in statistical inference and intermediate topics such as multiple least squares regression and ANOVA. The objective is to teach students to recognize situations where various statistical methods should be used, understand the strengths and limitations of the methods, and to show how they are implemented in R code. Examples are based on research described in the literature of behavioral ecology, with data sets and analysis code provided. Features: This intermediate to advanced statistical methods text was written with the behavioral ecologist in mind Computer programs are provided, written in the R language. Datasets are also provided, mostly based, at least to some degree, on real studies. Methods and ideas discussed include multiple regression and ANOVA, logistic and Poisson regression, machine learning and model identification, time-to-event modeling, time series and stochastic modeling, game-theoretic modeling, multivariate methods, study design/sample size, and what to do when things go wrong. It is assumed that the reader has already had exposure to statistics through a first introductory course at least, and also has sufficient knowledge of R. However, some introductory material is included to aid the less initiated reader. Scott Pardo, Ph.D., is an accredited professional statistician (PStat (R)) by the American Statistical Association. Michael Pardo is a Ph.D. is a candidate in behavioral ecology at Cornell University, specializing in animal communication and social behavior.

As experimental organisms, spiders offer ecologists a unique opportunity to examine the concept of the ecological community and the role which field experimentation can play in evaluating theories of population and community ecology. In this book, David Wise provides a balanced critique of field experiments designed to uncover details of spider ecology, with the dual aim of clarifying the ecology of these fascinating organisms and providing insight into the advantages and challenges of performing field experiments with a predator ubiquitous in terrestrial ecosystems. This book should be an essential reference for all ecologists wishing to learn more about the ecology of a major terrestrial predator and the use of field experimentation as a powerful technique to test ecological hypotheses.

The Heliconius butterflies are one of the classic systems in evolutionary biology and have contributed hugely to our understanding of evolution over the last 150 years. Their dramatic radiation and remarkable mimicry has fascinated biologists since the days of Bates, Wallace, and Darwin. The Ecology and Evolution of Heliconius Butterflies is the first thorough and accessible treatment of the ecology, genetics, and behaviour of these butterflies, exploring how they offer remarkable insights into tropical biodiversity. The book starts by outlining some of the evolutionary questions that Heliconius research has helped to address, then moves on to an overview of the butterflies themselves and their ecology and behaviour before focussing on wing pattern evolution, and finally, speciation. Richly illustrated with 32 colour plates, this book makes the extensive scientific literature on Heliconius butterflies accessible to a wide audience of professional ecologists, evolutionary biologists, entomologists, and amateur collectors.

Some of the most exciting recent advances in animal behaviour have occurred at the interface between that subject and the study of evolution. This book, written by experts in this area, illustrates how the profound changes in our understanding of evolution have influenced behavioural research. Its chapters span both studies of how behaviour itself has evolved, dealing with topics such as comparative studies, the genetics of behaviour, speciation and the evolution of sociality and of intelligence, and also the adaptiveness which this evolution has brought about, with treatment of mating and fighting strategies, and theories of kinship and altruism. The Evolution of Behaviour should be valuable to senior undergraduate and graduate students of biology and psychology, especially those studying animal behaviour, behavioural ecology, sociobiology, evolution, ecology and environmental biology.

Some of the most exciting recent advances in animal behaviour have occurred at the interface between that subject and the study of evolution. This book, written by experts in this area, illustrates how the profound changes in our understanding of evolution have influenced behavioural research. Its chapters span both studies of how behaviour itself has evolved, dealing with topics such as comparative studies, the genetics of behaviour, speciation and the evolution of sociality and of intelligence, and also the adaptiveness which this evolution has brought about, with treatment of mating and fighting strategies, and theories of kinship and altruism. The Evolution of Behaviour should be valuable to senior undergraduate and graduate students of biology and psychology, especially those studying animal behaviour, behavioural ecology, sociobiology, evolution, ecology and environmental biology.

How is the staggering biodiversity of the parasitoid insects maintained? This book, first published in 1994, explores patterns in host-parasitoid interactions, including parasitoid community richness, the importance of parasitoids as mortality factors, and their impact on host densities as determined by the outcomes of parasitoid introductions for biological control. It documents general patterns using data sets generated from the global literature and evaluates potential underlying biological, ecological and evolutionary mechanisms. A theme running throughout the book is the importance of host refuges as a major constraint on host-parasitoid interactions. Much can be learnt from the analysis of broad patterns; a few simple rules can go a long way in explaining the major components of these interactions. This book will be an invaluable resource for researchers interested in community ecology, population biology, entomology and biological control.

Acridids (grasshoppers and locusts) can range from being rare curiosities to abundant menaces. Some are threatened with extinction and become subjects of intensive conservation efforts, while others are devastating pests and become the objects of massive control programmes. Even within a species, there are times when the animal is so abundant that its crushed masses cause the wheels of trains to skid (the Rocky Mountain grasshopper, Melanoplus spretus Walsh in western North America in the 1860s and I 870s), while at other times the animal is alarmingly scarce (the Rocky Mountain grasshopper went extinct in the early 1900s). Why are there these extremes in one insect family, and even in a single species? The NATO workshop examined this paradox and its implications for Environmental Security, which must address both the elements of land use (agricultural production and pest management) and conservation of biodiversity. The reconciliation of these objectives clearly demands a critical assessment of current knowledge and policies, identification of future research, and close working relationships among scientists. Insects can present two clear faces, as well as the intervening gradation. These extremes require us to respond in two ways: conservation of scarce species and suppression of abundant (harmful) species. But perhaps most important, these opposite poles also provide the opportunity for an exchange of information and insight.

Ecological Methods by the late T.R. E. Southwood and revised over the years by P. A. Henderson has developed into a classic reference work for the field biologist. It provides a handbook of ecological methods and analytical techniques pertinent to the study of animals, with an emphasis on non-microscopic animals in both terrestrial and aquatic environments. It remains unique in the breadth of the methods presented and in the depth of the literature cited, stretching right back to the earliest days of ecological research. The universal availability of R as an open source package has radically changed the way ecologists analyse their data. In response, Southwood's classic text has been thoroughly revised to be more relevant and useful to a new generation of ecologists, making the vast resource of R packages more readily available to the wider ecological community. By focusing on the use of R for data analysis, supported by worked examples, the book is now more accessible than previous editions to students requiring support and ideas for their projects. Southwood's Ecological Methods provides a crucial resource for both graduate students and research scientists in applied ecology, wildlife ecology, fisheries, agriculture, conservation biology, and habitat ecology. It will also be useful to the many professional ecologists, wildlife biologists, conservation biologists and practitioners requiring an authoritative overview of ecological methodology.

Insect overwintering is a fascinating process involving many physiological, epidemiological, biochemical and behavioural changes. The study of the overwintering process can offer an insight into the development of insects, as well as help us to predict the patterns of crop damage and insect-borne disease caused by some insect species. This book provides a comprehensive account of the various forms of insect overwintering and highlights important areas of economic interest. It will be essential reading for advanced students and researchers in the fields of zoology, agriculture, forestry and ecology.

This book describes the Kluane Boreal Forest Ecosystem Project which operated from 1986 to 1996 in the southwestern Yukon. It begins by describing the area and its physical setting, and then the background of the project and the wisdom that had accumulated to 1986, on how this system might operate. The details of the experiments set up are presented, partly to help the reader appreciate the difficulty of working at -40 degrees and partly to aid the reader should they contemplate doing similar experiements in the future. Then they examine the three trophic levels of plants, the herbivores, and the predators in detail to provide some surprises about how the individual species operate within the overall system. Finally, they synthesize their findings in a model of the boreal forest vertebrate community, and provide an overview of what they have discovered and what remains to be done. Over the ten years of this project a large number of students and researchers have joined together to produce a picture which makes major advances in our understanding of the boreal forest ecosystem.

In this 1992 book, John Kennedy's point is that explicit anthropomorphism was well-nigh killed by fierce criticism from the radical Behaviourists, but that we have to recognize that today there is a new anthropomorphism which is much harder to avoid because it is unintended and largely unconscious. For that reason even those who if they were asked would firmly reject anthropomorphism nevertheless unwittingly slip into it from time to time. This book contains nineteen essays on behavioural concepts which have seldom been identified as anthropomorphic but in fact bear that connotation and lead to mistakes. Some of these, such as search images in birds and the learning of grammatical language by apes, have been seen through as errors after a time. A greater number, such as efference copy, goal-directedness, cognition and suffering in animals, are still current though not yet regarded as erroneous. The final chapter outlines things we can do to minimise the damage it does to the causal analysis of animal behaviour.

A problem every student of animal behaviour has to come to terms with is anthropomorphism, which means assuming that animals think or feel more or less as we do. Because of fierce criticism from behaviourists, explicit anthropomorphism is no longer scientifically respectable. John Kennedy's thesis is that anthropomorphism is not necessarily dead, but is lurking under different disguises. In fact, it still affects research, but is often unintended and therefore it goes unrecognized. He provides ample documentary evidence of the way researchers unconsciously slip into anthropomorphism. The book contains nineteen essays on behavioural concepts that have seldom been identified as anthropomorphic, but in fact bear that connotation and lead to mistakes. Some of these, such as search images in birds and the learning of grammatical language by apes, have been seen as errors after a time. A greater number, such as efference copy, goal-directedness, cognition, and suffering in animals, are still current though not yet regarded as erroneous. We can hardly hope to cure ourselves altogether of thinking anthropomorphically, and it can be very useful as a metaphor.

Conservation medicine focuses on the cross-over between ecosystem, animal and human health. Conservation biology emerged as a "crisis" discipline in the 1980s at the interface between ecology, environmental policy and management; and work in the biomedical and veterinary sciences is now being folded into conservation biology, to explore the connections between animal and human health. It traces the environmental sources of pathogens and pollutants in order to develop a rounded, interdisciplinary understanding of the ecological causes of changes in human and animal health, and the consequences of diseases to populations and ecological communities.

The two volumes of John Wiens' Ecology of Bird Communities, first published in 1992, are recognised as having applications and importance beyond the study of birds to the wider study of ecology in general. The books contain a detailed synthesis of our understanding of the patterns of organisation of bird communities and of the factors that may determine them, drawing from studies from all over the world. The author, however, does more than simply review findings in bird community ecology. By emphasizing how proper logic and methods have or have not been followed and how different viewpoints have developed historically and have led to controversy, he extends the scope of these books far beyond the study of birds. Volume 1 Foundations and Patterns explores why avian community ecologists ask the questions they do and what philosophical and methodological approaches they have used to answer such questions. Most of the book is devoted to a critical evaluation of what is known about the nature and organisation of bird communities.

The two volumes of John Wiens' Ecology of Bird Communities have applications and importance to the whole field of ecology. The books contain a detailed synthesis of our current understanding of the patterns of organisation of bird communities and of the factors that may determine them, drawing from studies from all over the world. By emphasizing how proper logic and methods have or have not been followed and how different viewpoints have developed historically and have led to controversy, the scope of these books are extended far beyond the study of birds. Processes and Variations discusses the way in which bird community patterns have been interpreted. This second volume examines how the complexity and variability of natural environments may influence efforts to discern and understand the nature of these communities. Graduate students and professionals in avian biology and ecology will find these volumes a valuable stimulus and guide to future field studies and theory development.

Our highly seasonal world restricts insect activity to brief portions of the year. This feature necessitates a sophisticated interpretation of seasonal changes and enactment of mechanisms for bringing development to a halt and then reinitiating it when the inimical season is past. The dormant state of diapause serves to bridge the unfavourable seasons, and its timing provides a powerful mechanism for synchronizing insect development. This book explores how seasonal signals are monitored and used by insects to enact specific molecular pathways that generate the diapause phenotype. The broad perspective offered here scales from the ecological to the molecular and thus provides a comprehensive view of this exciting and vibrant research field, offering insights on topics ranging from pest management, evolution, speciation, climate change and disease transmission, to human health, as well as analogies with other forms of invertebrate dormancy and mammalian hibernation.

Melanism: Evolution in Action describes investigations into a ubiquitous biological phenomenon, the existence of dark, or melanic, forms of many species of mammals, insects, and some plants. Melanism is a particularly exciting phenomenon in terms of our understanding of evolution. Unlike many other polymorphisms, the rise of a melanic population within a species is a visible alteration. Not only this, but melanism may sometimes occur dramatically quickly compared to other evolutionary change. Examples of melanism include one of the most famous illustrations of Darwinian natural selection, the peppered moth. This book, the first written on melanism since 1973, gives a lucid and up-to-date appraisal of the subject. The book is divided into ten chapters. The first four chapters place melanism into its historical and scientific context, with illustrations of its occurrence, and physical and genetic properties. Chapters 5-9 look in more detail at melanism in moths and ladybirds, explaining the diversity of evolutionary reasons for melanism, and the complexities underlying this apparently simple phenomenon. The final chapter shows how the study of melanism has contibuted to our understanding of biological evolution as a whole. Written in an engaging and readable style, by an author whose enthusiasm and depth of knowledge is apparent throughout, this book will be welcomed by all students and researchers in the fields of evolution, ecology, entomology, and genetics. It will also be of relevance to professional and amateur entomologists and lepidopterists alike.

The result of compiling widely scattered research on fish in tropical rivers, lakes and seas. A comprehensive overview of the ecology of fish communities in freshwater as well as marine environments.

Coral reef communities are among the most complex, mature and productive ecosystems on earth. Their activity resulted in the creation of vast lime constructions. Being extremely productive and having the function of a powerful biofilter, coral reefs play an important role in global biogeochemical processes and in the reproduction of food resources in tropical marine regions. All aspects of coral reef science are covered systematically and on the basis of a holistic ecosystem approach. The geological history of coral reefs, their geomorphology as well as biology including community structure of reef biota, their functional characteristics, physiological aspects, biogeochemical metabolism, energy balance, environmental problems and management of resources are treated in detail.

In the last ten years, the "comparative method" has been revolutionized by modern statistical ways of incorporating phylogenies into the design and analysis of comparative studies. The results of this revolution are particularly important in the study of animal behavior, which has relied on interspecific comparisons to infer universal trends and evolutionary patterns. The chapters of this edited volume consider the impact of modern phylogenetic comparative methods on the study of animal behavior and discuss the main issues that need to be considered in design and analysis of a comparative study, considers possible differences between the evolution of behavior and the evolution of morphology, and reviews how phylogenetic comparative studies have been used in certain areas of behavioral research.

Biologists since Darwin have been intrigued and confounded by the complex issues involved in the evolution and ecology of the social behavior of insects. The self-sacrifice of sterile workers in ant colonies has been particularly difficult for evolutionary biologists to explain. In this important new book, Andrew Bourke and Nigel Franks not only present a detailed overview of the current state of scientific knowledge about social evolution in ants, but also show how studies on ants have contributed to an understanding of many fundamental topics in behavioral ecology and evolutionary biology.

One of the substantial contributions of "Social Evolution in Ants" is its clear explanation of kin selection theory and sex ratio theory and their applications to social evolution in insects. Working to dispel lingering skepticism about the validity of kin selection and, more broadly, of "selfish gene" theory, Bourke and Franks show how these ideas underpin the evolution of both cooperation and conflict within ant societies. In addition, using simple algebra, they provide detailed explanations of key mathematical models. Finally, the authors discuss two relatively little-known topics in ant social biology: life history strategy and mating systems.

This comprehensive, up-to-date, and well-referenced work will appeal to all researchers in social insect biology and to scholars and students in the fields of entomology, behavioral ecology, and evolution.

The ecological literature on marsupials is dominated by descriptive natural history, and there has hitherto been little attempt at either synthesis or evolutionary interpretation. This book attempts to provide such a synthesis, by drawing on both the descriptive data base and predictions from the burgeoning literature on behavioural and evolutionary ecology. It documents the excellent potential the study of marsupials provides for resolution of theoretical questions of general importance in biology. It does this in three ways. First, by describing the impressive diversity of marsupial life history strategies and trophic roles. Second, by careful comparison with the eutherians, the scope of the marsupial radiation is used to analyse the role of developmental constraints and adaptive radiation in determining the diversification of higher taxa. Lastly, it is suggested that the accessibility of marsupial young during their obligatory pouch life facilitates measurement, manipulation and assessment of kinship not possible in other mammalian groups. Further special topics include marsupial/plant mutualism, marsupial competition and the empirical uses of mammals with simple life histories.