This book presents a major summary of what is currently known about North American amphibians and fossils in the Pleistocene. The only book available on the subject, it includes a general treatment of the Pleistocene and its effects on the physical and biological world, and features an illustrated bestiary of the herpetological families and genera as well as a discussion of the identification of fossils. The changes in the North American herpetofauna from the Cretaceous to the Pleistocene are cogently summarized, and complete herpetofaunal lists from the seven designated Pleistocene regions of North America are included. The concluding chapter details range adjustments and extinction patterns, and contrasts the evolutionary "stasis" of the herpetofauna with the avian and mammalian fauna. The work will be considered an essential reference for paleontologists.

This book simulates a historical walk through nature, teaching readers about the biodiversity on Earth in various eras with a focus on past terrestrial environments. Geared towards a student audience, using simple terms and avoiding long complex explanations, the book discusses the plants and animals that lived on land, the evolution of natural systems, and how these biological systems changed over time in geological and paleontological contexts. With easy-to-understand and scientifically accurate and up-to-date information, readers will be guided through major biological events from the Earth's past. The topics in the book represent a broad paleoenvironmental spectrum of interests and educational modules, allowing for virtual visits to rich geological times. Eras and events that are discussed include, but are not limited to, the much varied Quaternary environments, the evolution of plants and animals during the Cenozoic, the rise of angiosperms, vertebrate evolution and ecosystems in the Mesozoic, the Permian mass extinction, the late Paleozoic glaciation, and the origin of the first trees and land plants in the Devonian-Ordovician. With state-of-the art expert scientific instruction on these topics and up-to-date and scientifically accurate illustrations, this book can serve as an international course for students, teachers, and other interested individuals.

Studies of Sr isotopic composition of thousands of samples of marine sediments and fossils have yielded a curve of 87Sr/86Sr versus age for seawater Sr that extends back to 1 billion years. The ratio has fluctuated with large amplitude during this time period, and because the ratio is always uniform in the oceans globally at any one time, it is useful as a stratigraphic correlation and age-dating tool. The ratio also appears to reflect major tectonic and climatic events in Earth history and hence provides clues as to the causes, timing, and consequences of those events. The seawater 87Sr/86Sr ratio is generally high during periods marked by continent-continent collisions, and lower when continental topography is subdued, and seafloor generation rates are high. There is evidence that major shifts in the seawater ratio can be ascribed to specific orogenic events and correlate with large shifts in global climate.

Imaging and visualizing fossils in three dimensions with tomography is a powerful approach in paleontology. Here, the authors introduce select destructive and non-destructive tomographic techniques that are routinely applied to fossils and review how this work has improved our understanding of the anatomy, function, taphonomy, and phylogeny of fossil echinoderms. Building on this, this Element discusses how new imaging and computational methods have great promise for addressing long-standing paleobiological questions. Future efforts to improve the accessibility of the data underlying this work will be key for realizing the potential of this virtual world of paleontology.

This new treatise is comprised of 24 papers covering virtually every recent development regarding fossil horses, rhinos, tapirs, and their extinct relatives. It includes materials currently available in no other reference, such as information on the oldest known rhino and the oldest known perissodactyl, as well as papers debunking myths about the evolution of horses, and strong evidence that hyraxes are not related to elephants. The summary chapter includes useful charts which show the relationships of all perissodactyl genera, their distribution in time and space, and a new classification of the order.

The study of echinoid evolution, diversity, and ecology has always suffered from the fact that they are represented by taxa showing widely differing architectural designs of their multi-plated skeletons, inhabiting a large range of marine paleoenvironments, which result in highly varying taphonomic biases dictating their presence and recognition. This Element addresses the taphonomy of echinoids and includes: a general introduction to the morphological features of echinoids that play a role in their preservation; a review of processes which play an important role in the differential preservation of both regular and irregular echinoids including predation and transport; a summary of taphonomic pathways included in actualistic studies for recent sea urchins and then reconstructed for fossil taxa; and finally, a case study of the variation of echinoid taphonomy across a shelf gradient using the rich Miocene echinoid fauna of Sardinia.

Molybdenum (Mo) is a widely used trace metal for investigating redox conditions. However, unanswered questions remain that concentration and bulk isotopic analysis cannot specially answer. Improvements can be made by combining new geochemical techniques to traditional methods of Mo analysis. In this Element, we propose a refinement of Mo geochemistry within aquatic systems, ancient rocks, and modern sediments through molecular geochemistry (systematically combining concentration, isotope ratio, elemental mapping, and speciation analyses). Specifically, to intermediate sulfide concentrations governing Mo behavior below the 'switch-point' and dominant sequestration pathways in low oxygen conditions. The aim of this work is to 1) aid and improve the breadth of Mo paleoproxy interpretations by considering Mo speciation and 2) address outstanding research gaps concerning Mo systematics (cycling, partitioning, sequestration, etc.). The Mo paleoproxy has potential to solve ever complex research questions. By using molecular geochemical recommendations, improved Mo paleoproxy interpretations and reconstruction can be achieved.

Problematic fossils--those groups of organisms that do not fit conveniently into any existing phylum--play a pivotal role in the reconstruction of the history of life, being in effect "early experiments." This lavishly illustrated volume provides careful analyses and descriptions--in anatomical, functional, and developmental terms--of most of the major problematic fossil taxa. Sixteen original papers, written by internationally recognized scholars, discuss the features that make these taxa problematic and that provide clues to their phylogenetic relationships. Since Precambrian groups have been well covered in the existing literature, Hoffman and Nitecki focus on Paleozoic, and especially Early Paleozoic organisms, although Precambrian biota are also discussed.

Macroevolutionary inference has historically been treated as a two-step process, involving the inference of a tree, and then inference of a macroevolutionary model using that tree. Newer models blend the two steps. These methods make more complete use of fossils than the previous generation of Bayesian phylogenetic models. They also involve many more parameters than prior models, including parameters about which empiricists may have little intuition. In this Element, we set forth a framework for fitting complex, hierarchical models. The authors ultimately fit and use a joint tree and diversification model to estimate a dated phylogeny of the Cincta (Echinodermata), a morphologically distinct group of Cambrian echinoderms that lack the fivefold radial symmetry characteristic of extant members of the phylum. Although the phylogeny of cinctans remains poorly supported in places, this Element shows how models of character change and diversification contribute to understanding patterns of phylogenetic relatedness and testing macroevolutionary hypotheses.

This Open Access book introduces readers to the regional geology of Hanggai, Xianxia and Chuancun, the area between China's northern Zhejiang Province and southern Anhui Province and explores the strata, magmatic rocks and tectonic structures in 1:50,000 scale geological maps. Based on studies of multiple stratigraphic divisions, the standard stratigraphic section of the upper Ordovician Hirnantian in the lower Yangtze region is established, revealing for the first time numerous "Burgess Shale-type" sponge fossils in Hirnantian strata and identifying 10 grapholite fossil belts and various fossil categories, including chitin, trilobites, gastropods, brachiopods, and cephalopods. Moreover, the book identifies for the first time Late Ordovician volcanic events in northern Zhejiang province. The work represents a major contribution to research on Paleozoic strata in the Lower Yangtze region, and sheds new light on understanding the Hirnantian glacial event and biological extinction event in South China by providing a high-precision time scale. In addition, the book opens an important avenue for future research on sponge evolution after the Cambrian life explosion. As such, it offers a unique and valuable asset for researchers and graduate students alike.

This volume combines 10 years of accomplished research at the Pilauco site. The studies are focused on a variety of scientific areas including geological, sedimentological, geomorphological and paleobotanical topics, as well as paleontology of vertebrata and invertebrata, micropaleontology, archaeology, biochemistry, taxonomy, taphonomy, astrophysics and the development of some particular touristic aspects. In 18 chapters a variety of authors describe the excavation and investigation of this unique location. The book presents Pilauco as an example for the natural laboratory which can be found in South America, a testing ground for many of the hypotheses regarding migrations of animals and humans. In this context the study of topics, such as the paleozoography, the role of megafauna species for the architecture of the forests, the animal extinctions or the early human settlements, is extremely important on a global scale. The Pilauco site features paleontological and archaeological evidences and is contemporaneous with the Monte Verde site (~ 15,000 cal. yr AP). It is located 100 km north from Monte Verde and lies within the Intermediate Depression in northwestern Chilean Patagonia. It was discovered by chance in 1986 and has been excavated and investigated since 2007.

Modern videography provides an ever-widening window into subsea echinoderm life with vast potential for new knowledge. Supported by video evidence throughout, this Element begins with time-lapse video made in 1983 on film, using an off-the-shelf camera, flash, and underwater housings. Although quality has now been significantly improved by digital imagery, films from over thirty years ago captured crinoid feeding behavior previously unknown and demonstrated a great potential to learn about many other aspects of their biology. This sequence is followed by several examples of recent digital videography from submersibles of deep-sea crinoids and remotely operated vehicles (ROVs) (stalked and unstalked), as well as close-up video of crinoids in aquaria. These recent studies enabled a new classification of crinoid arm postures, provided detailed views of food particle capture, and revealed a wide range of behaviors in taxa never before seen in life.

A study of the Ichthyostega genus of extinct animals This detailed fossil study takes readers back millions of years to the Devonian Period. The Devonian Tetrapod Ichthyostega is Number 40 within the Fossils and Strata series. The series offers monographs and memoirs in palaeontology and stratigraphy. The international Fossils and Strata features systematic and regional monographs with taxonomic descriptions. The series is owned by and published on behalf of The Lethaia Foundation with collaboration among the Scandinavian countries.

Recent advances in statistical approaches called phylogenetic comparative methods (PCMs) have provided paleontologists with a powerful set of analytical tools for investigating evolutionary tempo and mode in fossil lineages. However, attempts to integrate PCMs with fossil data often present workers with practical challenges or unfamiliar literature. This Element presents guides to the theory behind and the application of PCMs with fossil taxa. Based on an empirical dataset of Paleozoic crinoids, example analyses are presented to illustrate common applications of PCMs to fossil data, including investigating patterns of correlated trait evolution and macroevolutionary models of morphological change. The authors emphasize the importance of accounting for sources of uncertainty and discuss how to evaluate model fit and adequacy. Finally, the authors discuss several promising methods for modeling heterogeneous evolutionary dynamics with fossil phylogenies. Integrating phylogeny-based approaches with the fossil record provides a rigorous, quantitative perspective on understanding key patterns in the history of life.

No animal shakes the human consciousness quite like a bear, and few compare to the giant short-faced bears that stalked North America during the Pleistocene. Even among the mammoths and saber-toothed cats, they were a staggering sight: on all fours, the biggest would stare a six-foot person in the face and weigh close to a ton. On hind legs they towered more than ten feet, with jaws powerful enough to crush skulls and snap bones like twigs. The bears weren't invincible, however. Despite their size, they were swept off the planet in a mysterious wave of Ice Age extinctions more than ten thousand years ago, then mostly forgotten. Chasing the Ghost Bear is Mike Stark's journey into the bear's enigmatic story-its life, disappearance, and rediscovery-and those trying to piece it together today. An engaging guide through his intrepid search, Stark's story leads us from the La Brea Tar Pits in Los Angeles to a cornfield in Indiana, the far ends of the Arctic, the plains of Texas, and the swamps of Florida. Part natural history, part travelogue, and part meditation on extinction and loss, Chasing the Ghost Bear returns these magnificent beasts to their rightful place in our understanding of the world just an epoch past.

Cetaceans (whales, dolphins, and porpoises) have fascinated and bewildered humans throughout history. Their mammalian affinities have been long recognized, but exactly which group of terrestrial mammals they descend from has, until recently, remained in the dark. Recent decades have produced a flurry of new fossil cetaceans, extending their fossil history to over 50 million years ago. Along with new insights from genetics and developmental studies, these discoveries have helped to clarify the place of cetaceans among mammals, and enriched our understanding of their unique adaptations for feeding, locomotion and sensory systems. Their continuously improving fossil record and successive transformation into highly specialized marine mammals have made cetaceans a textbook case of evolution - as iconic in its own way as the origin of birds from dinosaurs. This book aims to summarize our current understanding of cetacean evolution for the serious student and interested amateur using photographs, drawings, charts and illustrations.

Illustrating the concepts, methods, and applications of computer-assisted virtual reconstruction

Virtual Reconstruction demonstrates how readers can transform organisms from physical reality into virtual reality. Specifically, the authors provide the knowledge and tools needed to enable researchers to reconstruct fragmented and distorted fossil specimens in 3-D images so that their function, biomechanics, developmental changes, and evolutionary modifications can be determined. Likewise, the authors demonstrate how the same 3-D techniques can be used to enhance medical diagnosis and permit detailed planning of surgical intervention and forensic reconstruction on the basis of patient-specific anatomical data. They also provide a thorough introduction into modern methods of 3-D geometric-morphometric analysis.

All the steps of 3-D reconstruction are covered in detail, including data acquisition, processing, graphical representation, interactive manipulation, morphometric analysis, and rapid prototyping. Helping readers to bridge the gap between theory and practice, the authors have taken a four-pronged approach to their subject: The main text introduces basic concepts, provides technical information, presents practical applications, and discusses potential issues for further research Feature boxes offer schematic diagrams to simplify difficult concepts and case studies that demonstrate how concepts and methods are applied in real-world situations Appendices contain classical linear algebra formulation as well as standard data formats for text, standard images, biomedical images, and graphical objects Companion Internet site allows readers to experiment with sample data sets andapplets and provides links to tutorials, applets, databases, and technical definitions: www.wiley.com/go/virtualreconstruction

The two authors are internationally recognized as leaders and innovators in their field. Their text reflects more than a decade of research in computer-assisted paleoanthropology as well as their insightful lectures and courses in biomedical imaging, scientific visualization, and computational morphology.

This is the only reference that introduces readers, in the biosciences, to the concepts, methods, and applications of computer-assisted virtual reconstruction. It is an invaluable resource for researchers in physical anthropology, paleontology, morphology, anatomy, medicine, forensics, and primatology. Surgeons will find this book a reliable and state-of-the-art information source. Furthermore, the text is an excellent reference for computer scientists working in the biosciences.

The stable chromium (Cr) isotope system has emerged over the past decade as a new tool to track changes in the amount of oxygen in earth's ocean-atmosphere system. Much of the initial foundation for using Cr isotopes ( 53Cr) as a paleoredox proxy has required recent revision. However, the basic idea behind using Cr isotopes as redox tracers is straightforward-the largest isotope fractionations are redox-dependent and occur during partial reduction of Cr(VI). As such, Cr isotopic signatures can provide novel insights into Cr redox cycling in both marine and terrestrial settings. Critically, the Cr isotope system-unlike many other trace metal proxies-can respond to short-term redox perturbations (e.g., on timescales characteristic of Pleistocene glacial-interglacial cycles). The Cr isotope system can also be used to probe the earth's long-term atmospheric oxygenation, pointing towards low but likely dynamic oxygen levels for the majority of Earth's history.

The assassin's bullet misses, the Archduke's carriage moves forward, and a catastrophic war is avoided. So too with the history of life. Re-run the tape of life, as Stephen J. Gould claimed, and the outcome must be entirely different: an alien world, without humans and maybe not even intelligence. The history of life is littered with accidents and any twist or turn may lead to a completely different world. Now this view is challenged in Simon Conway Morris' exploration of the evidence demonstrating life's almost eerie ability to repeatedly navigate towards a single solution. Are all evolutionary inevitabilities limited to the suitability of a planet? Where are our counterparts across the galaxy? If the tape of life can only run on a suitable planet, it seems that such Earth-like planets are much rarer than hoped, and we remain inevitably humans in a lonely Universe. Simon Conway Morris is the Ad Hominen Professor in the Earth Science Department at the University of Cambridge. Morris is also a fellow of St. John's College and the Royal Society. His research focuses on the study of the constraints on evolution, and the historical processes that lead to the emergence of complexity, especially with respect to the construction of the major animal body plans in the Cambrian explosion. His work is central to palaeobiology, but is also of great interest to molecular biologists and bioastronomers. Previous published works include The Crucible of Creation: Burgess Shale and the Rise of Animals (Getty Center for Education in the Arts, 1999); and co-author of Solnhofen (Cambridge, 1990).

This book broadens the reader's knowledge base on lithofacies distribution, facies succession and association, and interpretation of paleo-depositional environments using outcrop-based and measured sedimentologic section data integrated with facies and petrographic analyses. Besides, the author also provides step-bystep workflow that could guide detailed geological field mapping and improve outcrop studies across Middle-Upper Cretaceous (Cenomanian-Campanian) successions of Southern Benue Trough and the lower stratigraphic interval of Anambra Basin, outcropping in Afikpo area of Southeastern Nigeria.

During the Early Cretaceous, lakes, meandering streams, and flood plains covered the region where the current foothills of Rioja now exist. Today the area is known for its wine and for the dozens of sites where footprints and trackways of dinosaurs, amphibians, and even pterosaurs can be seen. The dinosaurs that lived here 120 million years ago left their footsteps imprinted in the mud and moist soil. Now fossilized in rock, they have turned Rioja into one of the most valuable dinosaur footprint sites in all of Europe. Felix Perez-Lorente and his colleagues have published extensively on the region, mostly in Spanish-language journals. In this volume, Perez-Lorente provides an up-to-date synthesis of that research in English. He offers detailed descriptions of the sites, footprints, and trackways, and explains what these prints and tracks can tell us about the animals who made them."

The history of life on earth is largely reconstructed from time-averaged accumulations of fossils. A glimpse at ecologic-time attributes and processes is relatively rare. However, the time-sensitive and predictability of echinoderm disarticulation makes them model organisms to determine post-mortem transportation and allows recognition of ecological-time data within paleocommunity accumulations. Unlike many other fossil groups, this has allowed research on many aspects of echinoderms and their paleocommunities, such as the distribution of soft tissues, assessment of the amount of fossil transportation prior to burial, determination of intraspecific variation, paleocommunity composition, estimation of relative abundance of taxa in paleocommunities, determination of attributes of niche differentiation, etc. Crinoids and echinoids have received the most amount of taphonomic research, and the patterns present in these two groups can be used to develop a more thorough understanding of all echinoderm clades.

The quantification of morphology through time is a vital tool in elucidating macroevolutionary patterns. Studies of disparity require intense effort but can provide insights beyond those gained using other methodologies. Over the last several decades, studies of disparity have proliferated, often using echinoderms as a model organism. Echinoderms have been used to study the methodology of disparity analyses and potential biases as well as documenting the morphological patterns observed in clades through time. Combining morphological studies with phylogenetic analyses or other disparate data sets allows for the testing of detailed and far-reaching evolutionary hypotheses.