The study of echinoid evolution, diversity, and ecology has always suffered from the fact that they are represented by taxa showing widely differing architectural designs of their multi-plated skeletons, inhabiting a large range of marine paleoenvironments, which result in highly varying taphonomic biases dictating their presence and recognition. This Element addresses the taphonomy of echinoids and includes: a general introduction to the morphological features of echinoids that play a role in their preservation; a review of processes which play an important role in the differential preservation of both regular and irregular echinoids including predation and transport; a summary of taphonomic pathways included in actualistic studies for recent sea urchins and then reconstructed for fossil taxa; and finally, a case study of the variation of echinoid taphonomy across a shelf gradient using the rich Miocene echinoid fauna of Sardinia.

Molybdenum (Mo) is a widely used trace metal for investigating redox conditions. However, unanswered questions remain that concentration and bulk isotopic analysis cannot specially answer. Improvements can be made by combining new geochemical techniques to traditional methods of Mo analysis. In this Element, we propose a refinement of Mo geochemistry within aquatic systems, ancient rocks, and modern sediments through molecular geochemistry (systematically combining concentration, isotope ratio, elemental mapping, and speciation analyses). Specifically, to intermediate sulfide concentrations governing Mo behavior below the 'switch-point' and dominant sequestration pathways in low oxygen conditions. The aim of this work is to 1) aid and improve the breadth of Mo paleoproxy interpretations by considering Mo speciation and 2) address outstanding research gaps concerning Mo systematics (cycling, partitioning, sequestration, etc.). The Mo paleoproxy has potential to solve ever complex research questions. By using molecular geochemical recommendations, improved Mo paleoproxy interpretations and reconstruction can be achieved.

Problematic fossils--those groups of organisms that do not fit conveniently into any existing phylum--play a pivotal role in the reconstruction of the history of life, being in effect "early experiments." This lavishly illustrated volume provides careful analyses and descriptions--in anatomical, functional, and developmental terms--of most of the major problematic fossil taxa. Sixteen original papers, written by internationally recognized scholars, discuss the features that make these taxa problematic and that provide clues to their phylogenetic relationships. Since Precambrian groups have been well covered in the existing literature, Hoffman and Nitecki focus on Paleozoic, and especially Early Paleozoic organisms, although Precambrian biota are also discussed.

Macroevolutionary inference has historically been treated as a two-step process, involving the inference of a tree, and then inference of a macroevolutionary model using that tree. Newer models blend the two steps. These methods make more complete use of fossils than the previous generation of Bayesian phylogenetic models. They also involve many more parameters than prior models, including parameters about which empiricists may have little intuition. In this Element, we set forth a framework for fitting complex, hierarchical models. The authors ultimately fit and use a joint tree and diversification model to estimate a dated phylogeny of the Cincta (Echinodermata), a morphologically distinct group of Cambrian echinoderms that lack the fivefold radial symmetry characteristic of extant members of the phylum. Although the phylogeny of cinctans remains poorly supported in places, this Element shows how models of character change and diversification contribute to understanding patterns of phylogenetic relatedness and testing macroevolutionary hypotheses.

This Open Access book introduces readers to the regional geology of Hanggai, Xianxia and Chuancun, the area between China's northern Zhejiang Province and southern Anhui Province and explores the strata, magmatic rocks and tectonic structures in 1:50,000 scale geological maps. Based on studies of multiple stratigraphic divisions, the standard stratigraphic section of the upper Ordovician Hirnantian in the lower Yangtze region is established, revealing for the first time numerous "Burgess Shale-type" sponge fossils in Hirnantian strata and identifying 10 grapholite fossil belts and various fossil categories, including chitin, trilobites, gastropods, brachiopods, and cephalopods. Moreover, the book identifies for the first time Late Ordovician volcanic events in northern Zhejiang province. The work represents a major contribution to research on Paleozoic strata in the Lower Yangtze region, and sheds new light on understanding the Hirnantian glacial event and biological extinction event in South China by providing a high-precision time scale. In addition, the book opens an important avenue for future research on sponge evolution after the Cambrian life explosion. As such, it offers a unique and valuable asset for researchers and graduate students alike.

This volume combines 10 years of accomplished research at the Pilauco site. The studies are focused on a variety of scientific areas including geological, sedimentological, geomorphological and paleobotanical topics, as well as paleontology of vertebrata and invertebrata, micropaleontology, archaeology, biochemistry, taxonomy, taphonomy, astrophysics and the development of some particular touristic aspects. In 18 chapters a variety of authors describe the excavation and investigation of this unique location. The book presents Pilauco as an example for the natural laboratory which can be found in South America, a testing ground for many of the hypotheses regarding migrations of animals and humans. In this context the study of topics, such as the paleozoography, the role of megafauna species for the architecture of the forests, the animal extinctions or the early human settlements, is extremely important on a global scale. The Pilauco site features paleontological and archaeological evidences and is contemporaneous with the Monte Verde site (~ 15,000 cal. yr AP). It is located 100 km north from Monte Verde and lies within the Intermediate Depression in northwestern Chilean Patagonia. It was discovered by chance in 1986 and has been excavated and investigated since 2007.

Modern videography provides an ever-widening window into subsea echinoderm life with vast potential for new knowledge. Supported by video evidence throughout, this Element begins with time-lapse video made in 1983 on film, using an off-the-shelf camera, flash, and underwater housings. Although quality has now been significantly improved by digital imagery, films from over thirty years ago captured crinoid feeding behavior previously unknown and demonstrated a great potential to learn about many other aspects of their biology. This sequence is followed by several examples of recent digital videography from submersibles of deep-sea crinoids and remotely operated vehicles (ROVs) (stalked and unstalked), as well as close-up video of crinoids in aquaria. These recent studies enabled a new classification of crinoid arm postures, provided detailed views of food particle capture, and revealed a wide range of behaviors in taxa never before seen in life.

A study of the Ichthyostega genus of extinct animals This detailed fossil study takes readers back millions of years to the Devonian Period. The Devonian Tetrapod Ichthyostega is Number 40 within the Fossils and Strata series. The series offers monographs and memoirs in palaeontology and stratigraphy. The international Fossils and Strata features systematic and regional monographs with taxonomic descriptions. The series is owned by and published on behalf of The Lethaia Foundation with collaboration among the Scandinavian countries.

Recent advances in statistical approaches called phylogenetic comparative methods (PCMs) have provided paleontologists with a powerful set of analytical tools for investigating evolutionary tempo and mode in fossil lineages. However, attempts to integrate PCMs with fossil data often present workers with practical challenges or unfamiliar literature. This Element presents guides to the theory behind and the application of PCMs with fossil taxa. Based on an empirical dataset of Paleozoic crinoids, example analyses are presented to illustrate common applications of PCMs to fossil data, including investigating patterns of correlated trait evolution and macroevolutionary models of morphological change. The authors emphasize the importance of accounting for sources of uncertainty and discuss how to evaluate model fit and adequacy. Finally, the authors discuss several promising methods for modeling heterogeneous evolutionary dynamics with fossil phylogenies. Integrating phylogeny-based approaches with the fossil record provides a rigorous, quantitative perspective on understanding key patterns in the history of life.

Cetaceans (whales, dolphins, and porpoises) have fascinated and bewildered humans throughout history. Their mammalian affinities have been long recognized, but exactly which group of terrestrial mammals they descend from has, until recently, remained in the dark. Recent decades have produced a flurry of new fossil cetaceans, extending their fossil history to over 50 million years ago. Along with new insights from genetics and developmental studies, these discoveries have helped to clarify the place of cetaceans among mammals, and enriched our understanding of their unique adaptations for feeding, locomotion and sensory systems. Their continuously improving fossil record and successive transformation into highly specialized marine mammals have made cetaceans a textbook case of evolution - as iconic in its own way as the origin of birds from dinosaurs. This book aims to summarize our current understanding of cetacean evolution for the serious student and interested amateur using photographs, drawings, charts and illustrations.

Illustrating the concepts, methods, and applications of computer-assisted virtual reconstruction

Virtual Reconstruction demonstrates how readers can transform organisms from physical reality into virtual reality. Specifically, the authors provide the knowledge and tools needed to enable researchers to reconstruct fragmented and distorted fossil specimens in 3-D images so that their function, biomechanics, developmental changes, and evolutionary modifications can be determined. Likewise, the authors demonstrate how the same 3-D techniques can be used to enhance medical diagnosis and permit detailed planning of surgical intervention and forensic reconstruction on the basis of patient-specific anatomical data. They also provide a thorough introduction into modern methods of 3-D geometric-morphometric analysis.

All the steps of 3-D reconstruction are covered in detail, including data acquisition, processing, graphical representation, interactive manipulation, morphometric analysis, and rapid prototyping. Helping readers to bridge the gap between theory and practice, the authors have taken a four-pronged approach to their subject: The main text introduces basic concepts, provides technical information, presents practical applications, and discusses potential issues for further research Feature boxes offer schematic diagrams to simplify difficult concepts and case studies that demonstrate how concepts and methods are applied in real-world situations Appendices contain classical linear algebra formulation as well as standard data formats for text, standard images, biomedical images, and graphical objects Companion Internet site allows readers to experiment with sample data sets andapplets and provides links to tutorials, applets, databases, and technical definitions: www.wiley.com/go/virtualreconstruction

The two authors are internationally recognized as leaders and innovators in their field. Their text reflects more than a decade of research in computer-assisted paleoanthropology as well as their insightful lectures and courses in biomedical imaging, scientific visualization, and computational morphology.

This is the only reference that introduces readers, in the biosciences, to the concepts, methods, and applications of computer-assisted virtual reconstruction. It is an invaluable resource for researchers in physical anthropology, paleontology, morphology, anatomy, medicine, forensics, and primatology. Surgeons will find this book a reliable and state-of-the-art information source. Furthermore, the text is an excellent reference for computer scientists working in the biosciences.

The stable chromium (Cr) isotope system has emerged over the past decade as a new tool to track changes in the amount of oxygen in earth's ocean-atmosphere system. Much of the initial foundation for using Cr isotopes ( 53Cr) as a paleoredox proxy has required recent revision. However, the basic idea behind using Cr isotopes as redox tracers is straightforward-the largest isotope fractionations are redox-dependent and occur during partial reduction of Cr(VI). As such, Cr isotopic signatures can provide novel insights into Cr redox cycling in both marine and terrestrial settings. Critically, the Cr isotope system-unlike many other trace metal proxies-can respond to short-term redox perturbations (e.g., on timescales characteristic of Pleistocene glacial-interglacial cycles). The Cr isotope system can also be used to probe the earth's long-term atmospheric oxygenation, pointing towards low but likely dynamic oxygen levels for the majority of Earth's history.

This book broadens the reader's knowledge base on lithofacies distribution, facies succession and association, and interpretation of paleo-depositional environments using outcrop-based and measured sedimentologic section data integrated with facies and petrographic analyses. Besides, the author also provides step-bystep workflow that could guide detailed geological field mapping and improve outcrop studies across Middle-Upper Cretaceous (Cenomanian-Campanian) successions of Southern Benue Trough and the lower stratigraphic interval of Anambra Basin, outcropping in Afikpo area of Southeastern Nigeria.

During the Early Cretaceous, lakes, meandering streams, and flood plains covered the region where the current foothills of Rioja now exist. Today the area is known for its wine and for the dozens of sites where footprints and trackways of dinosaurs, amphibians, and even pterosaurs can be seen. The dinosaurs that lived here 120 million years ago left their footsteps imprinted in the mud and moist soil. Now fossilized in rock, they have turned Rioja into one of the most valuable dinosaur footprint sites in all of Europe. Felix Perez-Lorente and his colleagues have published extensively on the region, mostly in Spanish-language journals. In this volume, Perez-Lorente provides an up-to-date synthesis of that research in English. He offers detailed descriptions of the sites, footprints, and trackways, and explains what these prints and tracks can tell us about the animals who made them."

The history of life on earth is largely reconstructed from time-averaged accumulations of fossils. A glimpse at ecologic-time attributes and processes is relatively rare. However, the time-sensitive and predictability of echinoderm disarticulation makes them model organisms to determine post-mortem transportation and allows recognition of ecological-time data within paleocommunity accumulations. Unlike many other fossil groups, this has allowed research on many aspects of echinoderms and their paleocommunities, such as the distribution of soft tissues, assessment of the amount of fossil transportation prior to burial, determination of intraspecific variation, paleocommunity composition, estimation of relative abundance of taxa in paleocommunities, determination of attributes of niche differentiation, etc. Crinoids and echinoids have received the most amount of taphonomic research, and the patterns present in these two groups can be used to develop a more thorough understanding of all echinoderm clades.

The quantification of morphology through time is a vital tool in elucidating macroevolutionary patterns. Studies of disparity require intense effort but can provide insights beyond those gained using other methodologies. Over the last several decades, studies of disparity have proliferated, often using echinoderms as a model organism. Echinoderms have been used to study the methodology of disparity analyses and potential biases as well as documenting the morphological patterns observed in clades through time. Combining morphological studies with phylogenetic analyses or other disparate data sets allows for the testing of detailed and far-reaching evolutionary hypotheses.

Echinoderms elaborate a calcite skeleton composed of numerous plates with a distinct microstructure (stereom) that can be modelled into different shapes thanks to the use of a transient amorphous calcium carbonate (ACC) precursor phase and the incorporation of an intraorganic matrix during biomineralization. A variety of different types of stereom microarchitecture have been distinguished, each of them optimized for a specific function. For instance, a regular, galleried stereom typically houses collagenous ligaments, whereas an irregular, fine labyrinthic stereom commonly bears muscles. Epithelial tissues, in turn, are usually associated with coarse and dense stereom microfabrics. Stereom can be preserved in fossil echinoderms and a wide array of investigating methods are available. As many case studies have shown, a great deal of important paleobiological and paleoecological information can be decoded by studying the stereom microstructure of extinct echinoderms.

The principles of stratigraphic paleobiology can be readily applied to the nonmarine fossil record. Consistent spatial and temporal patterns of accommodation and sedimentation in sedimentary basins are an important control on stratigraphic architecture. Temperature and precipitation covary with elevation, causing significant variation in community composition, and changes in base level cause elevation to undergo predictable changes. These principles lead to eight sets of hypotheses about the nonmarine fossil record. Three relate to long-term and cyclical patterns in the preservation of major fossil groups and their taphonomy, as well as the occurrence of fossil concentrations. The remaining hypotheses relate to the widespread occurrence of elevation-correlated gradients in community composition, long-term and cyclical trends in these communities, and the stratigraphic position of abrupt changes in community composition. Testing of these hypotheses makes the stratigraphic paleobiology of nonmarine systems a promising area of investigation.

Arguably the best work to date in the history of geology.--David R. Oldroyd, Science After a superficial first glance, most readers of good will and broad knowledge might dismiss [this book] as being too much about too little. They would be making one of the biggest mistakes in their intellectual lives. . . . [It] could become one of our century's key documents in understanding science and its history.--Stephen Jay Gould, New York Review of Books Surely one of the most important studies in the history of science of recent years, and arguably the best work to date in the history of geology.--David R. Oldroyd, Science

This book discusses the Lagoa Santa Karst, which has been internationally known since the pioneering studies of the Danish naturalist Peter Lund in the early 1800s. It covers the speleogenesis, geology, vegetation, fauna, hydrogeology, geomorphology, and anthropogenic use of the Lagoa Santa Karst and is the first English-language book on this major karst area. The area, which has been at the heart of the debate on the origin and age of human colonization in the Americas, is characterized by a classical and scenic karst landscape with limestone cliffs, karst lakes and karst plains, in addition to numerous solution dolines. More than 1,000 caves have been documented in the area, many with significant archeological and paleontological value. Despite its great importance, the Lagoa Santa Karst faces severe environmental threats due to limestone mining and the expansion of the metropolis of Belo Horizonte and its surrounding towns. The growing recognition of the area's remarkable significance has led to increasing concern, and a number of protected areas have now been established, improving the conservation status of this landmark karst area.

Picture a world of dog-sized scorpions and millipedes as long as a car; tropical rainforests with trees towering over 150 feet into the sky and a giant polar continent five times larger than Antarctica. That world was not imaginary; it was the earth more than 300 million years ago in the Carboniferous period of the Paleozoic era. In Carboniferous Giants and Mass Extinction, George R. McGhee Jr. explores that ancient world, explaining its origins; its downfall in the end-Permian mass extinction, the greatest biodiversity crisis to occur since the evolution of animal life on Earth; and how its legacies still affect us today. McGhee investigates the consequences of the Late Paleozoic ice age in this comprehensive portrait of the effects of ancient climate change on global ecology. Carboniferous Giants and Mass Extinction examines the climatic conditions that allowed for the evolution of gigantic animals and the formation of the largest tropical rainforests ever to exist, which in time turned into the coal that made the industrial revolution possible-and fuels the engine of contemporary anthropogenic climate change. Exploring the strange and fascinating flora and fauna of the Late Paleozoic ice age world, McGhee focuses his analysis on the forces that brought this world to an abrupt and violent end. Synthesizing decades of research and new discoveries, this comprehensive book provides a wealth of insights into past and present extinction events and climate change.

Computational fluid dynamics (CFD), which involves using computers to simulate fluid flow, is emerging as a powerful approach for elucidating the palaeobiology of ancient organisms. Here, Imran A. Rahman describes its applications for studying fossil echinoderms. When properly configured, CFD simulations can be used to test functional hypotheses in extinct species, informing on aspects such as feeding and stability. They also show great promise for addressing ecological questions related to the interaction between organisms and their environment. CFD has the potential to become an important tool in echinoderm palaeobiology over the coming years.

The attraction of selenium isotopes as a paleoenvironmental tracer lies in the high redox potential of selenium oxyanions (SeIV and SeVI), the dominant species in the modern ocean. The largest isotopic fractionations occur during oxyanion reduction, which makes selenium isotopes a sensitive proxy for the redox evolution of our planet. As a case study we review existing data from the Neoarchean and Paleoproterozoic, which show that significant isotopic fractionations are absent until 2.5 Ga, and prolonged isotopic deviations only appear around 2.3 Ga. Selenium isotopes have thus begun to reveal complex spatiotemporal redox patterns not reflected in other proxies.

The diversity crisis in paleontology refers not to modern biota or the fossil record, but rather how our discipline lacks significant representation of individuals varying in race, ethnicity, and other aspects of identity. This Element is a call to action for broadening participation through improved classroom approaches as described in four sections. First, a brief review of the crisis and key concepts are presented. Next, culturally responsive pedagogy and related practices are introduced. Third, specific applications are offered for drawing cultural connections to studying the fossil record. Finally, recommendations including self-reflection are provided for fostering your own cultural competency. Our discipline offers much for understanding earth history and contributing new knowledge to a world impacted by humans. However, we must first more effectively welcome, support, and inspire all students to embrace meaning and value in paleontology; it is critical for securing the future of our field.