The first comprehensive synthesis of genomic techniques in earth sciences The past 15 years have witnessed an explosion of DNA sequencing technologies that provide unprecedented insights into biology. Although this technological revolution has been driven by the biomedical sciences, it also offers extraordinary opportunities in the earth and environmental sciences. In particular, the application of "omics" methods (genomics, transcriptomics, proteomics) directly to environmental samples offers exciting new vistas of complex microbial communities and their roles in environmental and geochemical processes. This unique book fills the gap where there exists a lack of resources and infrastructure to educate and train geoscientists about the opportunities, approaches, and analytical methods available in the application of omic technologies to problems in the geosciences. Genomic Approaches in Earth and Environmental Sciences begins by covering the role of microorganisms in earth and environmental processes. It then goes on to discuss how omics approaches provide new windows into geobiological processes. It delves into the DNA sequencing revolution and the impact that genomics has made on the geosciences. The book then discusses the methods used in the field, beginning with an overview of current technologies. After that it offers in-depth coverage of single cell genomics, metagenomics, metatranscriptomics, metaproteomics, and functional approaches, before finishing up with an outlook on the future of the field. The very first synthesis of an important new family of techniques Shows strengths and limitations (both practical and theoretical) of the techniques Deals with both theoretical and laboratory basics Shows use of techniques in a variety of applications, including various aspects of environmental science, geobiology, and evolution Genomic Approaches in Earth and Environmental Sciences is a welcome addition to the library of all earth and environmental scientists and students working within a wide range of subdisciplines.

One of the greatest mysteries in reconstructing the history of life on Earth has been the apparent absence of fossils dating back more than 550 million years. We have long known that fossils of sophisticated marine life-forms existed at the dawn of the Cambrian Period, but until recently scientists had found no traces of Precambrian fossils. The quest to find such traces began in earnest in the mid-1960s and culminated in one dramatic moment in 1993 when William Schopf identified fossilized microorganisms three and a half "billion" years old. This startling find opened up a vast period of time--some eighty-five percent of Earth's history--to new research and new ideas about life's beginnings. In this book, William Schopf, a pioneer of modern paleobiology, tells for the first time the exciting and fascinating story of the origins and earliest evolution of life and how that story has been unearthed.

Gracefully blending his personal story of discovery with the basics needed to understand the astonishing science he describes, Schopf has produced an introduction to paleobiology for the interested reader as well as a primer for beginning students in the field. He considers such questions as how did primitive bacteria, pond scum, evolve into the complex life-forms found at the beginning of the Cambrian Period? How do scientists identify ancient microbes and what do these tiny creatures tell us about the environment of the early Earth? (And, in a related chapter, Schopf discusses his role in the controversy that swirls around recent claims of fossils in the famed meteorite from Mars.) Like all great teachers, Schopf teaches the non-specialist enough about his subject along the way that we can easily follow his descriptions of the geology, biology, and chemistry behind these discoveries. Anyone interested in the intriguing questions of the origins of life on Earth and how those origins have been discovered will find this story the best place to start.

Dinosaurs are fascinating creatures and their popularity seems never ending, fuelled by films such as Jurassic Park and documentaries such as Walking with Dinosaurs. Yet dinosaurs (or more precisely non-avian dinosaurs) last trod the Earth 65 million years ago. All we know of them today are their fossilised bones, the tracks and traces that they left behind and, in very rare instances, some of the soft tissues or even traces of their chemistry. In many respects dinosaurs present us with one of the ultimate forensic challenges: they comprise the fragmentary remains of creatures that died many tens of millions of years ago, rather than just recently, or a few tens of years ago, which is the problem usually faced by forensic pathologists. How much do we really know about them, and to what extent can their remains inform us about ancient worlds, and indeed about the history of our planet? In this Very Short Introduction David Norman discusses how dinosaurs were first discovered and interpreted, and how our understanding of them has changed over the past 200 years. He looks at some of the amazing discoveries that have enabled us to gain new and unexpected insights into dinosaurs as animals with natural histories and behaviours, and considers some of the biggest questions in dinosaur biology, such as the implications of them having warm blood. Norman also shows how research upon dinosaurs has been enriched, particularly in recent decades, by technological break-throughs, which complement the informed speculation and luck which have played a part in many of the major discoveries. ABOUT THE SERIES: The Very Short Introductions series from Oxford University Press contains hundreds of titles in almost every subject area. These pocket-sized books are the perfect way to get ahead in a new subject quickly. Our expert authors combine facts, analysis, perspective, new ideas, and enthusiasm to make interesting and challenging topics highly readable.

This Open Access volume highlights how tree ring stable isotopes have been used to address a range of environmental issues from paleoclimatology to forest management, and anthropogenic impacts on forest growth. It will further evaluate weaknesses and strengths of isotope applications in tree rings. In contrast to older tree ring studies, which predominantly applied a pure statistical approach this book will focus on physiological mechanisms that influence isotopic signals and reflect environmental impacts. Focusing on connections between physiological responses and drivers of isotope variation will also clarify why environmental impacts are not linearly reflected in isotope ratios and tree ring widths. This volume will be of interest to any researcher and educator who uses tree rings (and other organic matter proxies) to reconstruct paleoclimate as well as to understand contemporary functional processes and anthropogenic influences on native ecosystems. The use of stable isotopes in biogeochemical studies has expanded greatly in recent years, making this volume a valuable resource to a growing and vibrant community of researchers.

'Vivid, thrilling, a delight ... Tim Flannery is a palaeontologist and ecologist of global standing, and this is a compelling and authoritative narrative of the evolution of Europe's flora and fauna, from the formation of the continent to its near future ... an exciting book, full of wonder' James McConnachie, Sunday Times A place of exceptional diversity, rapid change, and high energy, Europe has literally been at the crossroads of the world ever since the interaction of Asia, North America and Africa formed the tropical island archipelago that would become the continent of today. In this unprecedented evolutionary history, Tim Flannery shows how for the past 100 million years Europe has absorbed wave after wave of immigrant species; taking them in, transforming them, and sometimes hybridising them. Flannery reveals how, in addition to playing a vital role in the evolution of our own species, Europe was once the site of the formation of the first coral reefs, the home of some of the world's largest elephants, and now has more wolves than North America. This groundbreaking book charts the history of the land itself and the forces shaping life on it - including modern humans - to create a portrait of a continent that continues to exert a huge influence on the world today.

Fungi are ubiquitous in the world and responsible for driving the evolution and governing the sustainability of ecosystems now and in the past. Fossil Fungi is the first encyclopedic book devoted exclusively to fossil fungi and their activities through geologic time. The book begins with the historical context of research on fossil fungi (paleomycology), followed by how fungi are formed and studied as fossils, and their age. The next six chapters focus on the major lineages of fungi, arranging them in phylogenetic order and placing the fossils within a systematic framework. For each fossil the age and provenance are provided. Each chapter provides a detailed introduction to the living members of the group and a discussion of the fossils that are believed to belong in this group. The extensive bibliography (~ 2700 entries) includes papers on both extant and fossil fungi. Additional chapters include lichens, fungal spores, and the interactions of fungi with plants, animals, and the geosphere. The final chapter includes a discussion of fossil bacteria and other organisms that are fungal-like in appearance, and known from the fossil record. The book includes more than 475 illustrations, almost all in color, of fossil fungi, line drawings, and portraits of people, as well as a glossary of more than 700 mycological and paleontological terms that will be useful to both biologists and geoscientists.

Fossils have been vital to our understanding of the formation of the earth and the origins of all life on it. However, their impact has not been limited to debates about geology and evolution: attempts to explain their existence has shaken religion at its very roots, and they have remained a subject of ceaseless fascination for people of all ages and backgrounds. In this delightful book, Keith Thomson provides a remarkably all-encompassing explanation of fossils as a phenomenon. How did Darwin use fossils to support his theory of evolution? What are 'living fossils'? What fossils will we leave behind for future generations to examine? Building on the scientific aspects, he places fossils in a very human context, highlighting their impact on philosophy and mythology, our concept of time, and today's popular culture. What quickly becomes obvious is that the discovery of fossils and the ways in which they have been interpreted over time makes for fascinating reading. From the black market to the Piltdown Man, and from mythological dragons to living dinosaurs, fossils hold a permanent place in the popular imagination. ABOUT THE SERIES: The Very Short Introductions series from Oxford University Press contains hundreds of titles in almost every subject area. These pocket-sized books are the perfect way to get ahead in a new subject quickly. Our expert authors combine facts, analysis, perspective, new ideas, and enthusiasm to make interesting and challenging topics highly readable.

Land bridges are the causeways of biodiversity. When they form, organisms are introduced into a new patchwork of species and habitats, forever altering the ecosystems into which they flow; and when land bridges disappear or fracture, organisms are separated into reproductively isolated populations that can evolve independently. More than this, land bridges play a role in determining global climates through changes to moisture and heat transport and are also essential factors in the development of biogeographic patterns across geographically remote regions. In this book, paleobotanist Alan Graham traces the formation and disruption of key New World land bridges and describes the biotic, climatic, and biogeographic ramifications of these land masses' changing formations over time. Looking at five land bridges, he explores their present geographic setting and climate, modern vegetation, indigenous peoples (with special attention to their impact on past and present vegetation), and geologic history. From the great Panamanian isthmus to the boreal connections across the North Atlantic and North Pacific Oceans that allowed exchange of organisms between North America, Europe, and Asia, Graham's sweeping, one-hundred-million-year history offers new insight into the forces that shaped the life and land of the New World.

This thesis encompasses a study of past precipitation patterns based on six cave stalagmites from different parts of the Indian Himalaya. This is the first speleothem study in the Indian Himalaya that shows a direct relationship between past precipitation and the collapse of civilization. The stalagmites examined were KL-3 from Jammu and Kashmir; TCS and BR-1 from Himachal Pradesh; and DH-1, SA-1 and CH-1 from Uttarakhand. Based on the high-resolution palaeoclimatic reconstruction (35 U/th dates, 5 AMS dates, 1,500 samples for 18O and 13C values) obtained for the duration of the Pleistocene-Holocene transition (16.2-9.5 ka BP) and Mid-Holocene-Present (ca. 4.0 ka BP-Present), three major events were identified, namely the Older Dryas (OD), Bolling-Allerod (BA) period and Younger Dryas (YD) at ca. 14.3-13.9, 13.9-12.7 and 12.7-12.2 ka BP, respectively. The study showed a gradual reduction in the precipitation from 4 ka BP onwards for about a millennium with a peak arid period between 3.2 and 3.1 ka BP. According to the findings, the LIA (Little Ice Age) covers a time span from 1622-1820 AD, during which the climate was wetter than that in the post-LIA period (1820-1950 AD). In addition, this thesis supports the assumption that the WDs (Western Disturbances) contribute significantly to the total rainfall in the Himalaya region.

More than 99% of all life that has ever existed on this planet is extinct. Moreover, human acceleration of the extinction of species has created a crisis in biodiversity. How can the history of past life be retreived? How does this history bear on our understanding of the organization and evolution of present-day species? These questions are addressed in extinction and phylogeny.

A comprehensive pocket guide to dinosaurs and prehistoric animals and the world they inhabited millions of years ago. Packed with more than 700 full-colour illustrations, this definitive pocket guide paints a vivid portrait of extraordinary dinosaurs and prehistoric animals, and the ecosystems they lived in millions of years ago. Journey back in time and discover the incredible lives and habitats of dinosaurs, pterosaurs, marine reptiles, and prehistoric beasts. This fascinating guide explores every era of prehistoric life, from the Precambrian and Palaeozoic eras, when animals first began evolving to the Jurassic and Mesozoic era, which first saw the flourishing and then the eventual extinction of the dinosaurs. Learn more about prehistoric animals and the world they inhabited millions of years ago through detailed profiles of 200 dinosaurs and other ancestors of modern animals. These profiles are jargon-free and based on the latest science. This dinosaur encyclopedia is filled with crystal-clear artwork and photographs, fossils, and replica models showing the unique features of the different species. A Visual Guide to 200 Dinosaurs and Prehistoric Animals Part of the relaunched series of the popular DK Handbooks, this fact-filled book features a refreshed design and updated content. Packed with jaw-dropping images, fascinating dinosaur facts and straight-forward explanations, it's the perfect gift for budding palaeontologists of all ages or geology students. Inside the pages of this dinosaur encyclopedia from DK Books, you'll discover: - Maps indicating the location of where each animal's fossils have been found - Illustrations detailing the size of each animal in comparison to humans - Detailed, jargon-free profiles of 200 species with fun facts for quick reference

In the follow up to Darwin's Lost World, Martin Brasier book introduces the quest for the missing history of life and the cell. Through a series of journeys it emerges that the modern plant cell is one of the most deeply puzzling and unlikely steps in the whole history of life. Decoding this puzzle is a great adventure that has mainly taken place over the last half century. Brasier puts the big questions into context through lively descriptions of his explorations around the world, from the Caribbean Sea and the Egyptian pyramids, to the shores of the great lakes in Canada, andto the reefs and deserts of Australia. Covering the period from 1 to 2 billion years ago - a period he once dubbed 'the boring billion' - he demonstrates how it in fact involved great evolutionary potential with the formation of the complex (eukaryotic) cell. Without this cell there would be nothing on Earth today except bacteria, and the formation of this cell was a fundamental turning point in the history of life on Earth. Weaving together several threads, Brasier emphasises the importance of single-celled forms to marine ecosystems; symbiosis and coral reefs; and the architecture and beauty of single-celled Foraminifera and what they tell us about evolution. From a master storyteller comes a vivid description of the earliest biological forms and a set of fascinating tales of travels and research.

In the 19th and early 20th centuries, North American and European governments generously funded the discoveries of such famous paleontologists and geologists as Henry de la Beche, William Buckland, Richard Owen, Thomas Hawkins, Edward Drinker Cope, O. C. Marsh, and Charles W. Gilmore. In Patrons of Paleontology, Jane Davidson explores the motivation behind this rush to fund exploration, arguing that eagerness to discover strategic resources like coal deposits was further fueled by patrons who had a genuine passion for paleontology and the fascinating creatures that were being unearthed. These early decades of government support shaped the way the discipline grew, creating practices and enabling discoveries that continue to affect paleontology today.

Advances in Quaternary Entomology addresses the science of fossil insects by demonstrating their immense contribution to our knowledge of the paleoenvironmental and climatological record of the past 2.6 million years. In this comprehensive survey of the field, Scott A. Elias recounts development of scholarship, reviews the fossil insect record from Quaternary deposits throughout the world, and points to rewarding areas for future research. The study of Quaternary entomology is becoming an important tool in understanding past environmental changes. Most insects are quite specific as to habitat requirements, and those in non-island environments have undergone almost no evolutionary change in the Quaternary period. We therefore can use their modern ecological requirements as a basis for interpreting what past environments must have been like.
Features:
* describes and identifies principal characteristics of fossil insect groups of the Quaternary period
* Ties Quaternary insect studies to the larger field of paleoecology
* offers global coverage of the subject with specific regional examples
* illustrates specific methods and procedures for conducting research in Quaternary Entomology
* offers unique insight into overlying trends and broader implications of Quaternary climate change based on insect life of the period

This book highlights the knowledge about landscapes and characteristics of the earliest hunter-gatherer lifeway in Southern Patagonia. It presents an analysis of the archaeological investigations carried out during three decades by an interdisciplinary team that involved archaeologists, anthropologists, paleontologists, geologists and specialists in pollen and diatoms. The database yielded was recovered from systematic survey and excavations from the Pleistocene and Holocene stratigraphic layers of the rockshelter known as AEP-1, Piedra Museo Locality, situated in the central plateau of Santa Cruz Province, Argentina. Piedra Museo is a unique place in the world of high academic interest with some of the earliest archaeological remains in the Americas. Researchers defined two strata and several Stratigraphic units in the site based on the sedimentological and pedological characteristics. The depositional zones contain archaeological remains that are interpreted as hunting events corresponding to two main different occasions in the human colonization of the region, and a third human occupation during the Middle Holocene. Last one occurred then of the massive rockshelter roof colapse. The faunal remains led to a new approach to the palaeoenvironmental evolution of this enclosed basin. This volume describes the management of lithic raw materials and social networks from first human occupation of the Patagonian region to territorial consolidation of hunter-gatherer societies.

Although it is generally accepted that the Arctic Ocean is a very sensitive and important region for changes in the global climate, this region is the last major physiographic province of the earth whose short-and long-term geological history is much less known in comparison to other ocean regions. This lack of knowledge is mainly caused by the major technological/logistic problems in reaching this harsh, ice-covered region with normal research vessels and in retrieving long and undisturbed sediment cores. During the the last about 20 years, however, several international and multidisciplinary ship expeditions, including the first scientific drilling on Lomonosov Ridge in 2004, a break-through in Arctic research, were carried out into the central Artic and its surrounding shelf seas. Results from these expeditions have greatly advanced our knowledge on Arctic Ocean paleoenvironments.
Published syntheses about the knowledge on Arctic Ocean geology, on the other hand, are based on data available prior to 1990. A comprehensive compilation of data on Arctic Ocean paleoenvironment and its short-and long-term variability based on the huge amount of new data including the ACEX drilling data, has not been available yet. With this book, presenting (1) detailed information on glacio-marine sedimentary processes and geological proxies used for paleoenvironmental reconstructions, and (2) detailed geological data on modern environments, Quaternary variability on different time scales as well as the long-term climate history during Mesozoic-Tertiary times, this gap in knowledge will be filled.
*Aimed at specialists and graduates
*Presents background research, recent developments, and future trends
*Written by a leading scholar and industry expert

The graptolites constitute one of the geologically most useful taxonomic groups of fossils for dating rock successions, understanding paleobiogeography and reconstructing plate tectonic configurations in the Lower Palaeozoic. Graptolites were largely planktic, marine organisms, and as one of the first groups that explored the expanses of the world s oceans are vital for understanding Palaeozoic ecology. They are the best and often the only fossil group for dating Lower Palaeozoic rock successions precisely. Thousands of taxa have been described from all over the planet and are used for a wide variety of geological and palaeontological (biological) research topics. The recent recognition of the modern pterobranch Rhabdopleura as a living benthic graptolite enables a much better understanding and interpretation of the fossil Graptolithina. In the decades since the latest edition of the Graptolite Treatise, the enormous increase of knowledge on this group of organisms has never been synthesised in a compelling and coherent way, and information is scattered in scientific publications and difficult to sort through. This volume provides an up-to-date insight into research on graptolites. Such research has advanced considerably with the use of new methods of investigation and documentation. SEM investigation and research on ultrastructure of the tubaria has made it possible to compare extant and extinct taxa in much more detail. Cladistic interpretation of graptolite taxonomy and evolution has advanced the understanding of this group of organisms considerably in the last two decades, and has highlighted their importance in our understanding of evolutionary processes. This book will show graptolites, including their modern, living relatives, in a quite new and fascinating light, and will demonstrate the impact that the group has had on the evolution of the modern marine ecosystem. This book is aimed not only at earth scientists but also at biologists, ecologists and oceanographers. It is a readable and comprehensible volume for students at the MSc level, while remaining accessible to undergraduates and non-specialists seeking up-to-date information about this fascinating topic in palaeobiology.

The present volume is the first in a series of two books dedicated to the paleoceanography of the Late Cenozoic ocean. The need for an updated synthesis on paleoceanographic science is urgent, owing to the huge and very diversified progress made in this domain during the last decade. In addition, no comprehensive monography still exists in this domain. This is quite incomprehensible in view of the contribution of paleoceanographic research to our present understanding of the dynamics of the climate-ocean system. The focus on the Late Cenozoic ocean responds to two constraints. Firstly, most quantitative methods, notably those based on micropaleontological approaches, cannot be used back in time beyond a few million years at most. Secondly, the last few million years, with their strong climate oscillations, show specific high frequency changes of the ocean with a relatively reduced influcence of tectonics. The first volume addresses quantitative methodologies to reconstruct the dynamics of the ocean andthe second, major aspects of the ocean system (thermohaline circulation, carbon cycle, productivity, sea level etc.) and will also present regional synthesis about the paleoceanography of major the oceanic basins. In both cases, the focus is the "open ocean" leaving aside nearshore processes that depend too much onlocal conditions. In this first volume, we have gathered up-to-date methodologies for the measurement and quantitative interpretation of tracers and proxies in deep sea sediments that allow reconstruction of a few key past-properties of the ocean( temperature, salinity, sea-ice cover, seasonal gradients, pH, ventilation, oceanic currents, thermohaline circulation, andpaleoproductivity). Chapters encompass physical methods (conventional grain-size studies, tomodensitometry, magnetic and mineralogical properties), most current biological proxies (planktic and benthic foraminifers, deep sea corals, diatoms, coccoliths, dinocysts and biomarkers) and key geochemical tracers (trace elements, stable isotopes, radiogenic isotopes, and U-series). Contributors to the book and members of the review panel are among the best scientists in their specialty. They represent major European and North American laboratories and thus provide a priori guarantees to the quality and updat of the entire book. Scientists and graduate students in paleoclimatology, paleoceanography, climate modeling, and undergraduate and graduate students in marine geology represent the target audience. This volume should be of interest for scientists involved in several international programs, such as those linked to the IPCC (IODP - Integrated Ocean Drilling Program; PAGES - Past Global Changes; IMAGES - Marine Global Changes; PMIP: Paleoclimate Intercomparison Project; several IGCP projects etc.), That is, all programs that require access to time series illustrating changes in the climate-ocean system.
* Presents updated techniques and methods in paleoceanography
* Reviews the state-of-the-art interpretation of proxies used for quantitative reconstruction of the climate-ocean system
* Acts as a supplement for undergraduate and graduate courses in paleoceanography and marine geology

The vertebrate skeleton is one of nature's most amazing feats. Composed of cartilage and bone, it forms the supportive structure for all the remaining aspects of our anatomy. Stripped of skin, we can see the body's fascinating underlying architecture. In this one-of-a-kind book, biologist and skeletal reconstructionist Steve Huskey lays bare the vertebrate skeleton, providing a guided tour of the nuanced differences among the many featured vertebrate species. Using skeletal preparations he has spent decades assembling, Huskey helps us understand why animals live the way they do. He shows us the jaw and fang structures that allow venomous snakes to both kill and consume their prey whole. We see that the eastern mole is built like a weightlifter, allowing it to "swim through soil." Startling images demonstrate that the odd-looking trumpetfish is built not for music but for suction, with a skull that expands to vacuum in its prey. The pages of The Skeleton Revealed illuminate not only the elegance of each skeleton, but also the natural history story each skeleton tells. Come along-let's take a voyage through the boneyard.

Humans are the only mammals to walk on two, rather than four, legs. From an evolutionary perspective, this is an illogical development, as it slows us down. But here we are, suggesting there must have been something tremendous to gain from bipedalism. First Steps takes our ordinary, everyday walking experience and reveals how unusual and extraordinary it truly is. The seven-million-year-long journey through the origins of upright walking shows how it was in fact a gateway to many of the other attributes that make us human-from our technological skills and sociality to our thirst for exploration. DeSilva uses early human evolution to explain the instinct that propels a crawling infant to toddle onto two feet, differences between how men and women tend to walk, physical costs of upright walking, including hernias, varicose veins and backache, and the challenges of childbirth imposed by a bipedal pelvis. And he theorises that upright walking may have laid the foundation for the traits of compassion, empathy and altruism that characterise our species today and helped us become the dominant species on this planet.

During the Early Cretaceous, lakes, meandering streams, and flood plains covered the region where the current foothills of Rioja now exist. Today the area is known for its wine and for the dozens of sites where footprints and trackways of dinosaurs, amphibians, and even pterosaurs can be seen. The dinosaurs that lived here 120 million years ago left their footsteps imprinted in the mud and moist soil. Now fossilized in rock, they have turned Rioja into one of the most valuable dinosaur footprint sites in all of Europe. Felix Perez-Lorente and his colleagues have published extensively on the region, mostly in Spanish-language journals. In this volume, Perez-Lorente provides an up-to-date synthesis of that research in English. He offers detailed descriptions of the sites, footprints, and trackways, and explains what these prints and tracks can tell us about the animals who made them."

This book serves as an up-to-date introduction, as well as overview to modern trace fossil research and covers nearly all of the essential aspects of modern ichnology. Divided into three section, Trace Fossils covers the historical background and concepts of ichnology, on-going research problems, and indications about the possible future growth of the discipline and potential connections to other fields. This work is intended for a broad audience of geological and biological scientists. Workers new to the field could get a sense of the main concepts of ichnology and a clear idea of how trace fossil research is conducted. Scientists in related disciplines could find potential uses for trace fossils in their fields. And, established workers could use the book to check on the progress of their particular brand of ichnology. By design, there is something here for novice and veteran, insider and outsider, and for the biologically-oriented workers and for the sedimentary geologists.
* Presents a review of the state of ichnology at the beginning of the 21st Century
* Summarizes the basic concepts and methods of modern trace fossil research
* Discusses crucial background information about the history of trace fossil research, the main concepts of ichnology, examples of current problems and future directions, and the potential connections to other disciplines within both biology and geology