Although diversity is one of the central themes of ecology there is considerable disagreement ab out how it should be measured. I first encountered this problem 10 ycars ago whcn I started my research career and spent a long time pouring ovcr the literature in order to find the most useful techniques. The intervening decade has seen a further increasc in the number ofpapers devoted to the topic of ecological diversity but has led to no consensus on how it should be measured. My aim in writing this book is therefore to provide a practical guide to ecological diversity and its measurement. In a quantitative subject such as the measurement of diversity it is inevitable that some mathematics are involved, but at all times these are kept as simple as possible, and the emphasis is constantly on ecological reality and practical application. I hope that others ente ring thc fascinating ficld of ecological diversity will find it hclpful. This book grew out of my work in The School of Biological and Environmental Studies at the New University ofUlster, Coleraine, Northern Ircland. I am indebted to all the ecologists there for pro vi ding a stimulating atmosphere. Foremost among these were Amyan Macfadyen and Palmer Newbould. A number of the figures and tables in the book are based on data collected in Northern Irish woodlands.

This book celebrates the guppy's unique contribution to evolutionary ecology. Ever since Caryl Haskins described guppy populations as a 'natural experiment' because of the way predation pressure varies over a small geographical area, generations of researchers have been drawn to Trinidad to investigate evolution in the wild. The species continues to provide classic examples of natural selection in action and elegantly illustrates how ecology, evolution, and behaviour are interlinked.
Anne Magurran's account of the evolutionary ecology of the guppy integrates historical breakthroughs with new research in this fast-moving field. She reveals how guppies provided some of the first evidence of sperm competition and sexual selection, and how they continue to inform scientific thought on mating systems and cryptic choice. The consequences of variation in predation risk--as well as a host of other biotic and abiotic factors--are described and evaluated at all life stages from conception to death. The book discusses behavioural responses to ecological conditions alongside life history patterns. It examines the potential for ecological speciation and discusses new research into how reproductive isolating mechanisms become established in promiscuous mating systems. Conservation issues are also considered, both in terms of protecting the irreplaceable Trinidadian guppy system and in the context of invasion ecology.
This timely synthesis of research into a species that has raised key questions in evolutionary ecology will be of great interest to graduate level students as well as professional researchers in the fields of behavioural ecology and evolutionary biology.

Ecological diversity, or the variety and abundance of species in different habitats and communities, is one of the central themes of ecology. However, much of the existing literature on this subject is diffuse, often confusing, and in many cases complicated by unnecessarily difficult mathematics. This book aims to provide a succinct and clear summary of the relevant literature and a practical guide to the measurement of diversity.

The author discusses the methods of describing ecological diversity in conjunction with specific recommendations for the selection and interpretation of diversity measures. In addition, she considers the sampling problems often encountered in ecological censusing. The work concludes with a discussion of the empirical value of diversity measures. A special feature that makes the book particularly accessible to readers without great expertise in mathematics is the inclusion of worked examples of the main diversity measures and models.

Biological Diversity provides an up to date, authoritative review of the methods of measuring and assessing biological diversity, together with their application. The book's emphasis is on quantifying the variety, abundance, and occurrence of taxa, and on providing objective and clear guidance for both scientists and managers. This is a fast-moving field and one that is the focus of intense research interest. However the rapid development of new methods, the inconsistent and sometimes confusing application of old ones, and the lack of consensus in the literature about the best approach, means that there is a real need for a current synthesis. Biological Diversity covers fundamental measurement issues such as sampling, re-examines familiar diversity metrics (including species richness, diversity statistics, and estimates of spatial and temporal turnover), discusses species abundance distributions and how best to fit them, explores species occurrence and the spatial structure of biodiversity, and investigates alternative approaches used to assess trait, phylogenetic, and genetic diversity. The final section of the book turns to a selection of contemporary challenges such as measuring microbial diversity, evaluating the impact of disturbance, assessing biodiversity in managed landscapes, measuring diversity in the imperfect fossil record, and using species density estimates in management and conservation.

Biological Diversity provides an up to date, authoritative review of the methods of measuring and assessing biological diversity, together with their application. The book's emphasis is on quantifying the variety, abundance, and occurrence of taxa, and on providing objective and clear guidance for both scientists and managers. This is a fast-moving field and one that is the focus of intense research interest. However the rapid development of new methods, the inconsistent and sometimes confusing application of old ones, and the lack of consensus in the literature about the best approach, means that there is a real need for a current synthesis. Biological Diversity covers fundamental measurement issues such as sampling, re-examines familiar diversity metrics (including species richness, diversity statistics, and estimates of spatial and temporal turnover), discusses species abundance distributions and how best to fit them, explores species occurrence and the spatial structure of biodiversity, and investigates alternative approaches used to assess trait, phylogenetic, and genetic diversity. The final section of the book turns to a selection of contemporary challenges such as measuring microbial diversity, evaluating the impact of disturbance, assessing biodiversity in managed landscapes, measuring diversity in the imperfect fossil record, and using species density estimates in management and conservation.