When it comes to reproduction, gymnosperms are deeply weird. Cycads and co- fers have drawn out reproduction: at least 13 genera take over a year from polli- tion to fertilization. Since they don't apparently have any selection mechanism by which to discriminate among pollen tubes prior to fertilization, it is natural to w- der why such a delay in reproduction is necessary. Claire Williams' book celebrates such oddities of conifer reproduction. She has written a book that turns the context of many of these reproductive quirks into deeper questions concerning evolution. The origins of some of these questions can be traced back Wilhelm Hofmeister's 1851 book, which detailed the revolutionary idea of alternation of generations. This alternation between diploid and haploid generations was eventually to become one of the key unifying ideas in plant evolution. Dr. Williams points out that alter- tion of generations in conifers shows strong divergence in the evolution of male and female gametes, as well as in the synchronicity of male and female gamete development. How are these coordinated to achieve fertilization? Books on conifer reproduction are all too rare. The only major work in the last generation was Hardev Singh's 1978 Embryology of Gymnosperms, a book that summarized the previous century's work. Being a book primarily about embry- ogy, it stopped short of putting conifer reproduction in a genetic or evolutionary context.

What is the future of genetically modified (or transgenic) conifer plantations? The content of this edited volume Landscapes, Genomics and Transgenic Conifers addresses this question directly - and indirectly - using language drawn from policy, forest history, genomics, metabolism, pollen dispersal and gene flow, landscape ecology, evolution, economics, technology transfer and regulatory oversight. Although the book takes its title from a Nicholas School Leadership forum held November 17-19, 2004 at Duke University, its de novo contents move past the forum s deliberations. The result is a trans-disciplinary book composed of 14 chapters written by a total of 31 authors working in North America, South America, Europe and Africa.

The book is written for policy experts, life scientists, government and business leaders, biotechnology writers and activists. Few decision-makers realize the unprecedented degree to which transgenic technology is now possible for forests on a commercial scale. Only a handful of the 550 living conifer species is used for commodity value and even fewer species are being developed for transgenic plantations. Transgenic field trials started within the last decade but no transgenic pine plantations exist in 2005. But emergence of transgenic forest trees is still so recent that dialogue about the pros and cons is confined to the scientific community. And dialogue must move out into the public domain.

So little opportunity remains for opening public dialogue. The pursuit of transgenic research for forest trees is principally corporate so novel forest tree phenotypes are created as a means to increase shareholder value for investor companies. And while potential benefits will accrue to shareholders, ecological risks for certain transgenic traits are likely to be shared due to long-distance gene flow and inadequate bioconfinement measures. So this is a question riddled with tension. Without public deliberation, we should expect alienation of several interest groups. Alienation will lead to heightened clashes in the public policy arena or even radical environmental action. But how to move dialogue on transgenic forests forward? One must re-frame the issues behind transgenic conifer plantations.

The goal of this volume is to provide content for public deliberations about the genetic composition of future forests. Its Section I is composed of provocative and opposing views on the question of transgenic conifer plantations. Sections II and III follow with research advances on relevant conifer genomics and ecology research, respectively. Section IV forecasts rates of technology adoption for different case studies. Finally, Section V compares the status of regulatory oversight of transgenic forest trees between Canada and the United States. But will the book fulfil its goal? The burden of the answer lies with its readers. Will readers act or will transgenic forests be seen as too remote or simply too rural to bother with the angst of public deliberation? "

When it comes to reproduction, gymnosperms are deeply weird. Cycads and co- fers have drawn out reproduction: at least 13 genera take over a year from polli- tion to fertilization. Since they don't apparently have any selection mechanism by which to discriminate among pollen tubes prior to fertilization, it is natural to w- der why such a delay in reproduction is necessary. Claire Williams' book celebrates such oddities of conifer reproduction. She has written a book that turns the context of many of these reproductive quirks into deeper questions concerning evolution. The origins of some of these questions can be traced back Wilhelm Hofmeister's 1851 book, which detailed the revolutionary idea of alternation of generations. This alternation between diploid and haploid generations was eventually to become one of the key unifying ideas in plant evolution. Dr. Williams points out that alter- tion of generations in conifers shows strong divergence in the evolution of male and female gametes, as well as in the synchronicity of male and female gamete development. How are these coordinated to achieve fertilization? Books on conifer reproduction are all too rare. The only major work in the last generation was Hardev Singh's 1978 Embryology of Gymnosperms, a book that summarized the previous century's work. Being a book primarily about embry- ogy, it stopped short of putting conifer reproduction in a genetic or evolutionary context.