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No other disjunct pieces of land present such striking similarities
as the widely sepa 1 rated regions with a mediterranean type of
climate, that is, the territories fringing the Mediterranean Sea,
California, Central Chile and the southernmost strips of South
Mrica and Australia. Similarities are not confined to climatic
trends, but are also reflected in the physiognomy ofthe vegetation,
in land use patterns and frequently in the general appearance of
the landscape. The very close similarities in agricultural
practices and sometimes also in rural settlements are dependent on
the climatic and edaphic analogies, as well as on a certain
commonality in qdtural history. This is certainly true for the
Mediterranean Sea basin which in many ways represents a sort of
ecological-cultural unit; this is also valid for CaUfornia and
Chile, which were both settled by Spaniards and which showed
periods of vigorous commercial and cultural interchanges as during
the California gold rush. One other general feature is the massive
interchange of cultivated and weed species of plants that has
occurred between the five areas of the world that have a
mediterranean-type climate, with the Mediterranean basin region
itself as a major source. In spite of their limited territorial
extension, probably no other parts of the world have played a more
fundamental role in the history of mankind. Phoenician, Etruscan,
Hellenic, Jewish, Roman, Christian andArab civilizations, among
others, haveshapedmanyofman's present attitudes, including his
position and perception vis-a-vis nature."
The emergence of landscape ecology during the 1980s represents an
impor tant maturation of ecological theory. Once enamored with the
conceptual beauty of well-balanced, homogeneous ecosystems,
ecologists now assert that much of the essence of ecological
systems lies in their lumpiness. Patches with differing properties
and behaviors lie strewn across the land scape, products of the
complex interactions of climate, disturbance, and biotic processes.
It is the collective behavior of this patchwork of eco systems that
drives pattern and process of the landscape. is not an end point
This realization of the importance of patch dynamics in itself,
however. Rather, it is a passage to a new conceptual framework, the
internal workings of which remain obscure. The next tier of
questions includes: What are the fundamental pieces that compose a
landscape? How are these pieces bounded? To what extent do these
boundaries influence communication and interaction among patches of
the landscape? Will con sideration of the interactions among
landscape elements help us to under stand the workings of
landscapes? At the core of these questions lies the notion of the
ecotone, a term with a lineage that even predates ecosystem. Late
in the nineteenth century, F. E. Clements realized that the
transition zones between plant communi ties had properties distinct
from either of the adjacent communities. Not until the emergence of
patch dynamics theory, however, has central signif icance of the
ecotone concept become apparent."
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