Where We Stand Field workers-scientists of animal (including human!) behavior in nature-have long been fascinated by wild chimpanzees. A person who once has studied wild chimpanzees will be eager to observe them again. A person who has studied them twice will make every effort to continue the study, unless prevented from doing so. In short, behavioral primatology is addictive! Many people, among them Jane Goodall, Richard Wrangham, and I, do not regret that they have dedicated their whole lives to the study of wild chimpanzees. This is because the apes' behavior is always challenging: chimpanzees are cheerful, charming, playful, curious, beautiful, easygoing, generous, tolerant, and trustw- thy most of the time, but also are cautious, cunning, ugly, violent, ferocious, blo- thirsty, greedy, and disloyal at other times. We human beings share both the light and dark sides with our closest living relatives. For decades, we have documented huge across-population variation in behavior, as well as within-population variation. Cultural biology (now called cultural pri- tology), as proposed 60 years ago by Kinji Imanishi, recently has flourished.

The Great Apes (chimpanzees, bonobos, gorillas and orangutans) are our closest living relatives, sharing a common ancestor only five million years ago. We also share key features such as high intelligence, omnivorous diets, prolonged child-rearing and rich social lives. The Great Apes show a surprising diversity of adaptations, particularly in social life, ranging from the solitary life of orangutans, through patriarchy in gorillas to complex but different social organizations in bonobos and chimpanzees. As Great Apes are so close to humans, comparisons yield essential knowledge for modeling human evolutionary origins. Great Ape Societies provides comprehensive up-to-date syntheses of work on all four species, drawing on decades of international field work, zoo and laboratory studies. It will be essential reading for students and researchers in primatology, anthropology, psychology and human evolution.

The chimpanzee of all other living species is our closest relation, with whom we last shared a common ancestor about five million years ago. These African apes make and use a rich and varied kit of tools, and of the primates they are the only consistent and habitual tool-users and tool-makers. Chimpanzees meet the criteria of a culture as originally defined for human beings by socio-cultural anthropologists. They show sex differences in using tools to obtain and to process a variety of plant and animal foods. The technological gap between chimpanzees and human societies that live by foraging (hunter-gatherers) is surprisingly narrow at least for food-getting. Different communities of wild chimpanzees have different tool-kits and not all of this regional and local variation can be explained by the demands of the physical and biotic environments in which they live. Some differences are likely to be customs based on socially derived and symbolically encoded traditions. This book describes and analyzes the tool-use of humankind's nearest living relation. It focuses on field studies of these apes across Africa, comparing their customs to see if they can justifiably be termed cultural. It makes direct comparisons with the material culture of human foraging peoples. The book evaluates the chimpanzee as an evolutionary model, showing that chimpanzee behavior helps us to infer the origins of technology in human prehistory.

The chimpanzee of all other living species is our closest relation, with whom we last shared a common ancestor about five million years ago. These African apes make and use a rich and varied kit of tools, and of the primates they are the only consistent and habitual tool-users and tool-makers. Chimpanzees meet the criteria of a culture as originally defined for human beings by socio-cultural anthropologists. They show sex differences in using tools to obtain and to process a variety of plant and animal foods. The technological gap between chimpanzees and human societies that live by foraging (hunter-gatherers) is surprisingly narrow at least for food-getting. Different communities of wild chimpanzees have different tool-kits and not all of this regional and local variation can be explained by the demands of the physical and biotic environments in which they live. Some differences are likely to be customs based on socially derived and symbolically encoded traditions. This book describes and analyzes the tool-use of humankind's nearest living relation. It focuses on field studies of these apes across Africa, comparing their customs to see if they can justifiably be termed cultural. It makes direct comparisons with the material culture of human foraging peoples. The book evaluates the chimpanzee as an evolutionary model, showing that chimpanzee behavior helps us to infer the origins of technology in human prehistory.