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This book is the fourth in a series of 4 volumes in the Handbook of Zoology series about morphology, anatomy, reproduction, development, ecology, phylogeny and systematics of Annelida. It covers the most typical polychaetes, Phyllodocida, together with certain smaller taxa placed incertae sedis. This volume completes the polychaetous Annelida. Phyllodocida are often vagile, possess well-developed parapodia. Due to their broad and flat cirri these parapodia look like leaves in some taxa and leading to the name of the entire group. Many of its members are macrophagous and often predators. Accordingly most species possess elaborate sense structures such as sensory palps, antennae, eyes and nuchal organs. In certain species the eyes comprise thousands of photoreceptor cells and lenses most likely allowing forming true images. Phyllodocida typically possess an axial muscular pharynx called proboscis functioning as a kind of suction pipe allowing them to swallow and ingest their prey or other food. This pharynx may be armed with cuticular jaws and some species even possess venom glands. The probably most popular and important polychaete model organism, Platynereis dumerilii, belongs to this interesting group. Phyllodocida fall into two to three higher clades comprising about 25 families which represent more than one fourth of the polychaete diversity. One of these families, Syllidae, comprises about 700 valid species of mainly small size and may, therefore, represent one of the most complex and somehow difficult polychaete families on Earth.
This book is the second volume in a series of 4 volumes in the Handbook of Zoology series treating morphology, anatomy, reproduction, development, ecology, phylogeny, systematics and taxonomy of polychaetous Annelida. In this volume a comprehensive review of a few more derived higher taxa within Sedentaria are given, namely Sabellida, Opheliida/Capitellida as well as Hrabeiellidae. The former comprise annelids possessing a body divided into two more or less distinct regions or tagmata called thorax and abdomen. Here two groups of families are united, the spioniform and sabelliform polychaetes. Especially Spionidae and Sabellidae are speciose families within this group and represent two of the largest annelid families. These animals live in various types of burrows or tubes and all possess so-called feeding palps. In one group these appendages are differentiated as grooved feeding palps, whereas in the other they may form highly elaborated circular tentacular crowns comprising a number of radioles mostly giving off numerous filamentous pinnulae. Often additionally colourful, the latter are also received the common names "feather-duster worms", "flowers of the sea", "Christmas-tree worms". Opheliida/Capitellida including five families of truly worm-like annelids without appendages represents the contrary. Their members burrow in soft bottom substrates and may be classified as non-selective deposit feeders. Molecular phylogenetic analyses have shown that Echiura or spoon worms, formerly regarded to represent a separate phylum, are members of this group. Last not least Hrabeiellidae is one out of only two families of oligochaete-like terrestrial polychaetes and for this reason received strong scientific interest.
Annelida, mainly consisting of marine Polychaeta and in faunal and partly parasitic Clitellata, is one the most significant metazoan taxa. Its more than 20.000 described species invade nearly all habitats and play a central role in marine benthic systems as well as in terrestrial soil communities. Annelids include all soft-bodied segmented worm-like organisms and have been recognized as a separate "phylum" for almost 200 years. Recently, evidence has been accumulated which shows that some of the groups formerly regarded as independent "phyla" such as Pogonophora (now recognized as Siboglinidae), Echiura, Myzostomida and perhaps Sipuncula, are most probably nothing else than greatly modified Annelida. The extreme morphological diversity found especially in Polychaeta displays the plasticity of a simple segmented organisation that basically is nothing else but a serial repetition of identical units. Thus, annelids are highly important to our understanding of fundamental questions about morphological and adaptive diversity, as well as clarifying evolutionary changes and phylogenetic relationships. The book aims to summarize our knowledge on Polychaetes polychaetes and their allies and gives an overview of recent advances gained by studies that employed conventional and modern methods plus, increasingly and importantly, the use of molecular markers and computer-assisted kinship analyses. It also reflects the state of art in polychaete sciences and presents new questions and controversies. As such it will significantly influence the direction of research on Polychaeta and their related taxa.
This book is the first in a series of 4 volumes in the Handbook of Zoology series about morphology, anatomy, reproduction, development, ecology, phylogeny and systematics of Annelida. This first volume covers members of the so-called basal radiation and the first part of Sedentaria. It is supplemented by chapters on the history of annelid research, their fossil record, and an introduction to the phylogeny of annelids and their position in the tree of life. In the latter chapter the history of their systematic is reviewed giving an almost complete picture of systematic-scientific progress especially in the past years which changed our view on annelid phylogeny dramatically. The most basal annelids, lately united as Palaeoannelida, represent two families of aberrant polychaetes formerly often suggested to be highly derived which now give us a fresh look on how the ancestral annelid may have looked like. These lack certain key characters such as nuchal organs and possess rather simple nervous systems which now likely represent primitive character states. In this basal radiation the first taxon of apparently unsegmented and achaetigerous animals is positioned, the Sipuncula. Most likely another group of platyhelminth-like and unsegmented and even chaeta-lees annelids, Lobatocerebridae falls into this basal radiation. The section of Sedentaria starts with Orbiniida, a taxon characterized by elongated, thread-like worms which do not have anterior appendages like palps and comprises several families representing members of the Meiofauna. These minute worms often inhabiting the interstitial spaces in marine sands are suggested to have evolved by progenesis. The second higher taxon is represented by Cirratuliformia comprising nine families of typical sedentary polychaetes each of which showing a remarkable variation of the annelid body plan. Members of this taxon usually exhibit many annelid characters but certain also lack the most typical prostomial appendages, the palps.
Annelida, mainly consisting of marine Polychaeta and in faunal and partly parasitic Clitellata, is one the most significant metazoan taxa. Its more than 20.000 described species invade nearly all habitats and play a central role in marine benthic systems as well as in terrestrial soil communities. Annelids include all soft-bodied segmented worm-like organisms and have been recognized as a separate "phylum" for almost 200 years. Recently, evidence has been accumulated which shows that some of the groups formerly regarded as independent "phyla" such as Pogonophora (now recognized as Siboglinidae), Echiura, Myzostomida and perhaps Sipuncula, are most probably nothing else than greatly modified Annelida. The extreme morphological diversity found especially in Polychaeta displays the plasticity of a simple segmented organisation that basically is nothing else but a serial repetition of identical units. Thus, annelids are highly important to our understanding of fundamental questions about morphological and adaptive diversity, as well as clarifying evolutionary changes and phylogenetic relationships. The book aims to summarize our knowledge on Polychaetes polychaetes and their allies and gives an overview of recent advances gained by studies that employed conventional and modern methods plus, increasingly and importantly, the use of molecular markers and computer-assisted kinship analyses. It also reflects the state of art in polychaete sciences and presents new questions and controversies. As such it will significantly influence the direction of research on Polychaeta and their related taxa.
The nervous system is particularly fascinating for many biologists because it controls animal characteristics such as movement, behavior, and coordinated thinking. Invertebrate neurobiology has traditionally been studied in specific model organisms, whilst knowledge of the broad diversity of nervous system architecture and its evolution among metazoan animals has received less attention. This is the first major reference work in the field for 50 years, bringing together many leading evolutionary neurobiologists to review the most recent research on the structure of invertebrate nervous systems and provide a comprehensive and authoritative overview for a new generation of researchers. Presented in full colour throughout, Structure and Evolution of Invertebrate Nervous Systems synthesizes and illustrates the numerous new findings that have been made possible with light and electron microscopy. These include the recent introduction of new molecular and optical techniques such as immunohistochemical staining of neuron-specific antigens and fluorescence in-situ-hybridization, combined with visualization by confocal laser scanning microscopy. New approaches to analysing the structure of the nervous system are also included such as micro-computational tomography, cryo-soft X-ray tomography, and various 3-D visualization techniques. The book follows a systematic and phylogenetic structure, covering a broad range of taxa, interspersed with chapters focusing on selected topics in nervous system functioning which are presented as research highlights and perspectives. This comprehensive reference work will be an essential companion for graduate students and researchers alike in the fields of metazoan neurobiology, morphology, zoology, phylogeny and evolution.
Dieses konkurrenzlose Nachschlagewerk hat sich im Laufe der Jahre als zuverlassige Informationsquelle fur Zoologen und an Zoologie Interessierte erwiesen. Das nun in der 7. Auflage umfassend uberarbeitete Woerterbuch gibt in nunmehr 16.500 (!!) Stichwoertern prazise Auskunft uber alle wichtigen bzw. haufig vorkommenden Tiernamen und einschlagigen Fachwoerter. Es ist im deutschsprachigen Europa das einzige dieser Art und zeichnet sich aus durch hohe Informationsdichte, uberschaubare Gliederung und den Querschnittscharakter der aufgenommenen Stichwoerter.
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