What is animal welfare? Why has it proved so difficult to find a definition that everyone can agree on? This concise and accessible guide is for anyone who is interested in animals and who has wondered how we can assess their welfare scientifically. It defines animal welfare as 'health and animals having what they want', a definition that can be easily understood by scientists and non-scientists alike, expresses in simple words what underlies many existing definitions, and shows what evidence we need to collect to improve animal welfare in practice. Above all, it puts the animal's own point of view at the heart of an assessment of its welfare. But, can we really understand what animals want? A consistent theme running through the book is that not only is it possible to establish what animals want, but that this information is vital in helping us to make sense of the long and often confusing list of welfare measures that are now in use such as 'stress' and 'feel good hormones', expressive sounds and gestures, natural behaviour, cognitive bias, and stereotypies. Defining welfare as 'health and what animals want' allows us to distinguish between measures that are simply what an animal does when it is alert, aroused, or active and those measures that genuinely allow us to distinguish between situations the animals themselves see as positive or negative. Sentience (conscious feelings of pleasure, pain, and suffering) is for many people the essence of what is meant by welfare, but studying consciousness is notoriously difficult, particularly in non-human species. These difficulties are discussed in the context of our current - and as yet incomplete - knowledge of human and animal consciousness. Finally, the book highlights some key ideas in the relationship between animal welfare science and animal ethics and shows how closely the well-being of humans is linked to that of other animals. The Science of Animal Welfare is an ideal companion for undergraduate and graduate students taking courses in animal behaviour and welfare, as well as for professional researchers, practitioners and animal welfare consultants. At the same time, it is easily understandable to non-scientists and anyone without prior knowledge but with an interest in animals and the rapidly evolving science of animal welfare.

Evolutionary change is usually incremental and continuous, but some increases in organizational complexity have been radical and divisive. Evolutionary biologists, who refer to such events as "major transitions", have not always appreciated that these advances were novel forms of pairwise commitment that subjugated previously independent agents. Inclusive fitness theory convincingly explains cooperation and conflict in societies of animals and free-living cells, but to deserve its eminent status it should also capture how major transitions originated: from prokaryote cells to eukaryote cells, via differentiated multicellularity, to colonies with specialized queen and worker castes. As yet, no attempt has been made to apply inclusive fitness principles to the origins of these events. Domains and Major Transitions of Social Evolution develops the idea that major evolutionary transitions involved new levels of informational closure that moved beyond looser partnerships. Early neo-Darwinians understood this principle, but later social gradient thinking obscured the discontinuity of life's fundamental organizational transitions. The author argues that the major transitions required maximal kinship in simple ancestors - not conflict reduction in already elaborate societies. Reviewing more than a century of literature, he makes testable predictions, proposing that open societies and closed organisms require very different inclusive fitness explanations. It appears that only human ancestors lived in societies that were already complex before our major cultural transition occurred. We should therefore not impose the trajectory of our own social history on the rest of nature. This thought-provoking text is suitable for graduate-level students taking courses in evolutionary biology, behavioural ecology, organismal developmental biology, and evolutionary genetics, as well as professional researchers in these fields. It will also appeal to a broader, interdisciplinary audience, including the social sciences and humanities.

From foraging patterns in a single tree to social interactions across a home range, how primates use space is a key question in the field of primate behavioral ecology. Drawing on the latest advances in spatial analysis tools, this book offers practical guidance on applying geographic information systems (GIS) to central questions in primatology. An initial methodological section discusses niche modelling, home range analysis and agent-based modelling, with a focus on remote data collection. Research-based chapters demonstrate how ecologists apply this technology to a suite of topics including: calculating the intensity of use of both range and travel routes, assessing the impacts of logging, mining and hunting, and informing conservation strategies.

In a world increasingly concerned with climate change, food security, and other human issues, the welfare of non-human animals is in danger of being overlooked and side-lined. Using the latest scientific research on animal consciousness and emotions, Marian Stamp Dawkins argues that if animal welfare is to be taken seriously by world opinion, it needs a complete rethink. She asks important questions such as: are we justified in projecting human emotions on to animals? What can science tell us about their quality of life? She concludes that we need to place less emphasis on the conscious experience of suffering in animals, and more emphasis on the practical importance of animal welfare to human health and human well-being. This requires a long, hard look at some of the cherished ideas we hold about animal emotions, and what we can and cannot know about the conscious experiences of other animals.

This volume had it's inception in a series of eight lectures delivered at Columbia University during the spring of 1905, and represents, in a much condensed form, the results of a decade of uninterrupted study of the Formicidae, and of the works that have been written on these insects.

The more we learn about bees, the more extraordinary they seem. They have five eyes and beat their wings at almost 200 cycles per second. And to communicate with each other bees do make sounds but also dance. This book, packed with over 30 figures, gives a fascinating insight into the extraordinary complex and highly organised world of bees. The author, K. Weiss, explains the critical role that bees and bee-keeping play for human society, and offers advice for those interested in raising bees.

Crustaceans are increasingly used as model organisms in all fields of biology, including neurobiology, developmental biology, animal physiology, evolutionary ecology, biogeography, and resource management. One reason for the increasing use of crustacean examples is the wide range of phenotypes found in this group and the diversity of environments they inhabit; few other taxa exhibit such a variety of body shapes and adaptations to particular habitats and environmental conditions. A good overview of their functional morphology is essential to understanding many aspects of their biology. This volume is the first in The Natural History of Crustacea series, a ten-volume series that will treat all aspects of crustacean biology, physiology, behavior, and evolution. The series updates and synthesizes a growing wealth of information on the natural history of this remarkable group. Functional Morphology and Diversity explores the functional morphology of crustaceans, which cover the main body parts and systems. The book brings together a group of internationally recognized-and up-and-coming-experts in fields related to systematics and morphology. Contributing authors study a range of crustacean taxa and topics, and thus the volume provides a compact overview of the great phenotypic diversity and their function found among crustaceans. The first broad treatment of Crustacea in decades, the book will be invaluable for researchers and students in this and related fields.

How similar are your choices, behaviours, and lifestyle to those of a parrot? We humans are not like other mammals. We look like them, but we don't act like them. In fact, many of our defining human traits: our longevity, intelligence, monogamy and childrearing, and learning and language, all deep parts of what it means to be human, are far more similar to birds than to our fellow mammals. These similarities originate not from shared ancestors but from parallel histories. Our evolutionary stories have pushed humans and birds to the same solutions. In this book, Antone Martinho-Truswell explores these similarities to argue that we can learn a great deal about ourselves by thinking of the human species as 'the bird without feathers'. This is also a book about convergent evolution - evolution that drives very different species to very similar outcomes and behaviours. The traits we share with birds but not mammals are the result of similar, specific pressures that demanded similar solutions - and exploring these similarities can help us understand both why we evolved to be the way we are, and also how very unusual some of our behaviours are in the animal kingdom, Drawing on a rich array of examples across the natural world, Martinho-Truswell also demonstrates the ways in which parrots are our biological mirror image; an evolutionary parallel to ourselves. In contemplating what we share with the birds, and especially the parrots, we understand how close nature came to creating another lineage of radical intelligence on Earth, and we also come to better understand ourselves.

Anthrozoology, the study of human-animal interactions (HAIs), has experienced substantial growth during the past 20 years and it is now timely to synthesise what we know from empirical evidence about our relationships with both domesticated and wild animals. Two principal points of focus have become apparent in much of this research. One is the realisation that the strength of these attachments not only has emotional benefits for people, but confers health benefits as well, such that a whole area has opened up of using companion animals for therapeutic purposes. The other is the recognition that the interactions we have with animals have consequences for their welfare too, and thus impact on their quality of life. Consequently we now study HAIs in all scenarios in which animals come into contact with humans, whether as pets/companions, farm livestock, laboratory animals, animals in zoos, or in the wild. This topical area of study is of growing importance for animals in animal management, animal handling, animal welfare and applied ethology courses, and also for people within psychology, anthropology and human geography at both the undergraduate and postgraduate level. It will therefore be of interest to students, researchers, and animal managers across the whole spectrum of human-animal contact.

Why do baby sharks, hyenas, and pelicans kill their siblings? Why do beetles and mice commit infanticide? Why are twins and birth defects more common in older human mothers? "A Natural History of Families" concisely examines what behavioral ecologists have discovered about family dynamics and what these insights might tell us about human biology and behavior. Scott Forbes's engaging account describes an uneasy union among family members in which rivalry for resources often has dramatic and even fatal consequences.

In nature, parents invest resources and control the allocation of resources among their offspring to perpetuate their genetic lineage. Those families sometimes function as cooperative units, the nepotistic and loving havens we choose to identify with. In the natural world, however, dysfunctional familial behavior is disarmingly commonplace.

While explaining why infanticide, fratricide, and other seemingly antisocial behaviors are necessary, Forbes also uncovers several surprising applications to humans. Here the conflict begins in the moments following conception as embryos struggle to wrest control of pregnancy from the mother, and to wring more nourishment from her than she can spare, thus triggering morning sickness, diabetes, and high blood pressure. Mothers, in return, often spontaneously abort embryos with severe genetic defects, allowing for prenatal quality control of offspring.

Using a broad sweep of entertaining examples culled from the world of animals and humans, "A Natural History of Families" is a lively introduction to the behavioral ecology of the family.

When Charles Darwin first proposed Sexual Selection Theory, he suggested two mechanisms: competition among males and choice by females. Although their importance is long established and extremely well understood, their mirror images have remained largely underappreciated; males also choose, and females also compete. The combination of male mate choice (MMC) and female competition (FC) may be one of the most overlooked yet important and intriguing phenomena in modern sexual selection theory. This novel text reviews our current understanding of MMC and FC, highlighting the important connections between them. It places both concepts in the context of related fields such as female choice, mating systems, and sexual selection theory more broadly. A truly holistic approach is provided which takes all the relevant elements into consideration, especially the relative roles of MMC and FC, female ornamentation, their evolutionary consequences, and their genetic basis. Considering male mate choice and female competition in this way as effectively two sides of the same coin creates a powerful paradigm for a more complete understanding of sexual selection. Male Choice, Female Competition, and Female Ornaments in Sexual Selection will be suitable for both graduate students and researchers interested in sexual selection from an evolutionary, psychological, and anthropological perspective. It will also appeal to a broader audience of behavioural ecologists and evolutionary psychologists.

What is animal welfare? Why has it proved so difficult to find a definition that everyone can agree on? This concise and accessible guide is for anyone who is interested in animals and who has wondered how we can assess their welfare scientifically. It defines animal welfare as 'health and animals having what they want', a definition that can be easily understood by scientists and non-scientists alike, expresses in simple words what underlies many existing definitions, and shows what evidence we need to collect to improve animal welfare in practice. Above all, it puts the animal's own point of view at the heart of an assessment of its welfare. But, can we really understand what animals want? A consistent theme running through the book is that not only is it possible to establish what animals want, but that this information is vital in helping us to make sense of the long and often confusing list of welfare measures that are now in use such as 'stress' and 'feel good hormones', expressive sounds and gestures, natural behaviour, cognitive bias, and stereotypies. Defining welfare as 'health and what animals want' allows us to distinguish between measures that are simply what an animal does when it is alert, aroused, or active and those measures that genuinely allow us to distinguish between situations the animals themselves see as positive or negative. Sentience (conscious feelings of pleasure, pain, and suffering) is for many people the essence of what is meant by welfare, but studying consciousness is notoriously difficult, particularly in non-human species. These difficulties are discussed in the context of our current - and as yet incomplete - knowledge of human and animal consciousness. Finally, the book highlights some key ideas in the relationship between animal welfare science and animal ethics and shows how closely the well-being of humans is linked to that of other animals. The Science of Animal Welfare is an ideal companion for undergraduate and graduate students taking courses in animal behaviour and welfare, as well as for professional researchers, practitioners and animal welfare consultants. At the same time, it is easily understandable to non-scientists and anyone without prior knowledge but with an interest in animals and the rapidly evolving science of animal welfare.

Urban Evolutionary Biology fills an important knowledge gap on wild organismal evolution in the urban environment, whilst offering a novel exploration of the fast-growing new field of evolutionary research. The growing rate of urbanization and the maturation of urban study systems worldwide means interest in the urban environment as an agent of evolutionary change is rapidly increasing. We are presently witnessing the emergence of a new field of research in evolutionary biology. Despite its rapid global expansion, the urban environment has until now been a largely neglected study site among evolutionary biologists. With its conspicuously altered ecological dynamics, it stands in stark contrast to the natural environments traditionally used as cornerstones for evolutionary ecology research. Urbanization can offer a great range of new opportunities to test for rapid evolutionary processes as a consequence of human activity, both because of replicate contexts for hypothesis testing, but also because cities are characterized by an array of easily quantifiable environmental axes of variation and thus testable agents of selection. Thanks to a wide possible breadth of inference (in terms of taxa) that may be studied, and a great variety of analytical methods, urban evolution has the potential to stand at a fascinating multi-disciplinary crossroad, enriching the field of evolutionary biology with emergent yet incredibly potent new research themes where the urban habitat is key. Urban Evolutionary Biology is an advanced textbook suitable for graduate level students as well as professional researchers studying the genetics, evolutionary biology, and ecology of urban environments. It is also highly relevant to urban ecologists and urban wildlife practitioners.

Essential Ornithology provides the reader with a concise but comprehensive introduction to the biology of birds, one of the most widely studied taxonomic groups. The book begins by considering the dinosaur origins of birds and their subsequent evolution. Development, anatomy, and physiology are then discussed followed by chapters devoted to avian reproduction, migration, ecology, and conservation. Sections dealing with aspects of bird/human relationships and bird conservation give the book an applied context. This new edition has been thoroughly updated, providing new information from rapidly-developing fields including the avian fossil record, urban and agricultural ecology, responses to climate change, invasive species biology, technologies to track movement, avian disease, and the role of citizen scientists. There is also a greater focus on North American ornithology. Drawing extensively upon the wider scientific literature, this engaging text places the results of classical studies of avian biology alongside the most recent scientific breakthroughs. Useful case studies are presented in a concise and engaging style with the student reader foremost in mind. Key points are highlighted and suggestions for guided reading and key references are included throughout. Essential Ornithology is a companion textbook for advanced undergraduate and graduate students taking courses in avian science, as well as a useful reference for professional researchers and consultants. Amateur ornithologists will also find this book offers a scientifically rigorous and accessible overview for a more general readership.

Communicating with and understanding dogs builds trusting relationships, and relieves frustration and improves lives on both ends of the leash. Apart from simply reading canine body language, this book explores the juncture between hearts and minds where true understanding begins. It takes a light-hearted, yet earnest approach to exploring how to connect with dogs. Each chapter features amusing, thought-provoking anecdotes and illustrations that show us how dogs are our teachers, and how, by connecting with them, we learn how to connect with ourselves. This book helps us find the intersection between the academic and the spiritual for better communication with dogs. It's about having an effective two-way conversation; it shows us how to talk with our dogs instead of talking at them. The intuitive side of communicating with canines in conjunction with the intellectual side is also investigated: in this space, we build trust, respect, and better relationships, and human and canine lives are enriched. Come walk with us on our journey.

Primates communicate with each other using a wide range of signals: olfactory signals to mark territories, screams to recruit help while fighting, gestures to request food and facial expressions to initiate play. Primate Communication brings together research on all forms of interchange and discusses what we know about primate communication via vocal, gestural, facial, olfactory and integrated multimodal signals in relation to a number of central topics. It explores the morphological, neural and cognitive foundations of primate communication through discussion of cutting-edge research. By considering signals from multiple modalities and taking a unified multimodal approach, the authors offer a uniquely holistic overview of primate communication, discussing what we know, what we don't know and what we may currently misunderstand about communication across these different forms. It is essential reading for researchers interested in primate behaviour, communication and cognition, as well as students of primatology, psychology, anthropology and cognitive sciences.

All students and researchers of behaviour - from those observing freely-behaving animals in the field to those conducting more controlled laboratory studies - face the problem of deciding what exactly to measure. Without a scientific framework on which to base them, however, such decisions are often unsystematic and inconsistent. Providing a clear and defined starting point for any behavioural study, this is the first book to make available a set of principles for how to study the organisation of behaviour and, in turn, for how to use those insights to select what to measure. The authors provide enough theory to allow the reader to understand the derivation of the principles, and draw on numerous examples to demonstrate clearly how the principles can be applied. By providing a systematic framework for selecting what behaviour to measure, the book lays the foundations for a more scientific approach for the study of behaviour.

Widespread across open lands and cities of Europe, Africa, and Asia, the common kestrel (Falco tinnunculus) is one of the most abundant and studied birds of prey. This book brings together and synthesises the results of research on kestrels for professional ornithologists and scientists that seek to consolidate a vast body of literature. It is also a reference for those readers who may not have the depth of scientific knowledge to navigate new fields of scientific enquiry. It examines many aspects of the species' biology, from the reproductive strategies to the behavioural and demographic adaptations to changes of environmental conditions. It also discusses the roles of physiology and immunology in mediating the adaptability of kestrels to the ongoing environmental changes with a particular focus on contaminants. This volume presents new and exciting avenues of research on the ecology and behaviour of the common kestrel.

Few animals have a worse reputation than the vulture. But is it deserved? With Vulture, Katie Fallon offers an irresistible argument to the contrary, tracing a year in the life of a typical North American turkey vulture. Turkey vultures, also known as buzzards, are the most widely distributed and abundant scavenging birds of prey on the planet, found from central Canada to the southern tip of Argentina and nearly everywhere in between. Deftly drawing on the most up-to-date scientific papers and articles and weaving those in with interviews with world-renowned raptor and vulture experts and her own compelling natural history writing, Fallon examines all aspects of the bird’s natural history: breeding, incubating eggs, raising chicks, migrating, and roosting. The result is an intimate portrait of an underappreciated bird—one you’ll never look at in the same way again.

The essential textbook on agent-based modeling-now fully updated and expanded Agent-Based and Individual-Based Modeling has become the standard textbook on the subject for classroom use and self-instruction. Drawing on the latest version of NetLogo and fully updated with new examples, exercises, and an enhanced text for easier comprehension, this is the essential resource for anyone seeking to understand how the dynamics of biological, social, and other complex systems arise from the characteristics of the agents that make up these systems. Steven Railsback and Volker Grimm lead students stepwise through the processes of designing, programming, documenting, and doing scientific research with agent-based models, focusing on the adaptive behaviors that make these models necessary. They cover the fundamentals of modeling and model analysis, introduce key modeling concepts, and demonstrate how to implement them using NetLogo. They also address pattern-oriented modeling, an invaluable strategy for modeling real-world problems and developing theory. This accessible and authoritative book focuses on modeling as a tool for understanding real complex systems. It explains how to pose a specific question, use observations from actual systems to design models, write and test software, and more. A hands-on introduction that guides students from conceptual design to computer implementation to analysis Filled with new examples and exercises and compatible with the latest version of NetLogo Ideal for students and researchers across the natural and social sciences Written by two leading practitioners Supported by extensive instructional materials at www.railsback-grimm-abm-book.com

Animals have evolved remarkable biomechanical and physiological systems that enable their rich repertoire of motion. Animal Locomotion offers a fundamental understanding of animal movement through a broad comparative and integrative approach, including basic mathematics and physics, examination of new and enduring literature, consideration of classic and cutting-edge methods, and a strong emphasis on the core concepts that consistently ground the dizzying array of animal movements. Across scales and environments, this book integrates the biomechanics of animal movement with the physiology of animal energetics and the neural control of locomotion. This second edition has been thoroughly revised, incorporating new content on non-vertebrate animal locomotor systems, studies of animal locomotion that have inspired robotic designs, and a new chapter on the use of evolutionary approaches to locomotor mechanisms and performance.

While we joke that men are from Mars and women are from Venus, our gender differences can't compare to those of many other animals. For instance, the male garden spider spontaneously dies after mating with a female more than fifty times his size. And male blanket octopuses employ a copulatory arm longer than their own bodies to mate with females that outweigh them by four orders of magnitude. Why do these gender gulfs exist? Introducing readers to important discoveries in animal behavior and evolution, Odd Couples explores some of the most extraordinary sexual differences in the animal world. Daphne Fairbairn uncovers the unique and bizarre characteristics of these remarkable species and the special strategies they use to maximize reproductive success. Fairbairn also considers humans and explains that although we are keenly aware of our own sexual differences, they are unexceptional within the vast animal world. Looking at some of the most amazing creatures on the planet, Odd Couples sheds astonishing light on what it means to be male or female in the animal kingdom.

Why do dogs behave in the ways that they do? Why did our ancestors tame wolves? How have we ended up with so many breeds of dog, and how can we understand their role in contemporary human society? Explore the answers to these questions and many more in this study of the domestic dog. Building on the strengths of the first edition, this much-anticipated update incorporates two decades of new evidence and discoveries on dog evolution, behavior, training, and human interaction. It includes seven entirely new chapters covering topics such as behavioral modification and training, dog population management, the molecular evidence for dog domestication, canine behavioral genetics, cognition, and the impact of free-roaming dogs on wildlife conservation. It is an ideal volume for anyone interested in dogs and their evolution, behavior and ever-changing roles in society.