This book offers a unique collection of inter- and multidisciplinary studies on river systems. Rivers have been the prime source of sustenance since the advent of civilization and river systems often form the basis for agriculture, transport, water, and land for domestic, commercial, and industrial activities, fostering economic prosperity. A river basin is a basic geographical and climatological unit within which the vagaries of natural processes act and manifest themselves at different spatio-temporal scales. Even if compared side-by-side, no two river basins respond to natural processes in the same way and thus, it has long been recognized that each river basin is unique. Hence, any developmental activity or conservation effort has to be designed and implemented to match each unique river basin. With the burgeoning population and increasing dependency on natural resources, understanding and maintaining river systems has become increasingly important. This book provides a varied reference work on and unprecedented guidelines for conducting and implementing research on river basins, and for managing their ecological development.

4 Water Sources ........................................ 149 Criteria ............................................. 149 Major types .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 150 . . . . . . . . . . . . Summary ............................................ 152 5 Water Treatment ...................................... 155 Requirements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 155 . . . . . . . . . . . . Materials ............................................ 155 Treatment options . . . . . . . . . . . . . . . . . . . . . . . . . . . 156 . . . . . . . . . . . System design ........................................ 169 System monitoring and control . . . . . . . . . . . . . . . . . . . . . 172 . . . . . . . . Environmental considerations .............................. 174 Summary ............................................ 174 6 Culture Units ......................................... 175 Considerations in choosing culture units ...................... 175 Characteristics of culture units . . . . . . . . . . . . . . . . . . . . . 175 . . . . . . . . Applications of culture units .............................. 191 Hatchery design " . . . . . . . . . . . . . . . . . . . . . . . . . . 208 . . . . . . . . . . . Summary ............................................ 210 7 Obtaining Fish for Stocking . ............................. 211 Stock from the wild .................................... 211 Stock from the hatchery ................................. 211 Spermatogenesis (sperm formation) ....................... 232 Oogenesis (egg formation) ............................. 232 Oocyte maturation ................................... 233 Endocrine control of oocyte maturation and ovulation .......... 237 fuduced ovulation . . . . . . . . . . . . . . . . . . . . . . . . . . 238 . . . . . . . . . . Timing and egg quality . . . . . . . . . . . . . . . . . . . . . . . 257 . . . . . . . . . Artificial fertilization ................................. 265 Care of eggs ....................................... 267 Storage of gametes ................. ' .................. 269 Natural ovulation . . . . . . . . . . . . . . . . . . . . . . . . . . 270 . . . . . . . . . . Care of broodfish . . . . . . . . . . . . . . . . . . . . . . . . . . 289 . . . . . . . . . . Egg collection .. . . . . . . . . . . . . . . . . . . . . . . . . . . 290 . . . . . . . . . . fuduced vs natural ovulation ............................ 290 Broodfish adaptability . . . . . . . . . . . . . . . . . . . . . . . . . 291 . . . . . . . . . . Examples ............................................ 291 Genetic considerations . . . . . . . . . . . . . . . . . . . . . . . . . 295 . . . . . . . . . . Hybridization . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 296 . . . . . . . . . . . . Sex control .......................................... 296 Summary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 298 . . . . . . . . . . . . . vi 8 Nutrition of Larval Fish . . . . . . . . . . . . . . . . . . . . . . . 299 . . . . . . . . . . Feeding criteria ....................................... 299 Choice and culture of foods . . . . . . . . . . . . . . . . . . . . . . 307 . . . . . . . . . General feeding practices . . . . . . . . . . . . . . . . . . . . . . . 336 . . . . . . . . . . Specific feeding practices ................................ 352 General methods used in our hatchery . . . . . . . . . . . . . . . . . 372 . . . . . . . Industrial-scale larval food processing in Italian hatcheries ......... 373 Summary ............................................ 374 9 Nutrition of Juvenile and Adult Fish ...................... 375 ............................. 375 Requirements and components Broodstock nutrition .................................... 407 Nutritional disorders .................................... 408 Environmental considerations . . . . . . . . . . . . . . . . . . . . . 411 . . . . . . . . . Feed studies . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 411 . . . . . . . . . . . . Suggested feed formulas ................................. 460 Making and storing feeds ................................ 461 Feeding methods ...................................... 464 Summary ............................................ 467 10 Energetics ............................................ 469 Energy budget components and influencing factors . . . . . . . . . . . 469 . . . .

Echinoderms are an ancient and diverse group of marine animals with a rich fossil record. They occur abundantly in all modern oceans and at all depths, where they contribute importantly to patterns in biodiversity and to the structure and functioning of marine systems. It is therefore vital to understand how they will respond to a rapidly changing ocean climate and other anthropogenic stressors, informed by both the dynamics of the fossil record and responses of extant species. The theme of the 13th International Echinoderm Conference (Hobart, Tasmania, 5-9 January 2009) was the response of echinoderms to global change. Echinoderms in a Changing World contains a selection of plenary and contributed papers, and a comprehensive presentation of abstracts of all oral papers and posters. The collection will be useful to all students of echinoderm biology, ecology and palaeontology, from undergraduate level to professional researchers.

Selenium is a naturally occurring trace element that can become concentrated and released by industrial, agricultural, petrochemical and mining activities. At concentrated levels it is toxic and has polluted ecosystems around the world. This book will serve as a comprehensive practical handbook for everyone dealing with selenium in aquatic environments. It offers field-tested approaches and methods for assessment and water quality management. Using his twenty-year experience, the author discusses the effects of selenium on fish and bird populations and presents guidelines for identifying sources of pollution, interpreting selenium concentrations, assessing hazardous conditions, setting water quality criteria and ecosystem loading limits (TMDLs). He also includes a procedure for setting environmentally safe limits that ensure compliance with EPA regulations. Selenium Assessment in Aquatic Ecosystems will interest field scientists, natural resource managers, risk assessors and environmental planners.

Biological invasions are considered to be one of the greatest threats to the integrity of most ecosystems on earth. This volume explores the current state of marine bioinvasions, which have been growing at an exponential rate over recent decades. Focusing on the ecological aspects of biological invasions, it elucidates the different stages of an invasion process, starting with uptake and transport, through inoculation, establishment and finally integration into new ecosystems. Basic ecological concepts - all in the context of bioinvasions - are covered, such as propagule pressure, species interactions, phenotypic plasticity, and the importance of biodiversity. The authors approach bioinvasions as hazards to the integrity of natural communities, but also as a tool for better understanding fundamental ecological processes. Important aspects of managing marine bioinvasions are also discussed, as are many informative case studies from around the world.

Who was Nicolas Rashevsky? To answer that question, this book draws on Rashevsky's unexplored personal archival papers and shares interviews with his family, students and friends, as well as discussions with biologists and mathematical biologists, to flesh out and complete the picture. "Most modern-day biologists have never heard of Rashevsky. Why?" In what constitutes the first detailed biography of theoretical physicist Nicolas Rashevsky (1899-1972), spanning key aspects of his long scientific career, the book captures Rashevsky's ways of thinking about the place mathematical biology should have in biology and his personal struggle for the acceptance of his views. It brings to light the tension between mathematicians, theoretical physicists and biologists when it comes to the introduction of physico-mathematical tools into biology. Rashevsky's successes and failures in his efforts to establish mathematical biology as a subfield of biology provide an important test case for understanding the role of theory (in particular mathematics) in understanding the natural world. With the biological sciences moving towards new vistas of inter- and multi-disciplinary collaborations and research programs, the book will appeal to a wide readership ranging from historians, sociologists, and ethnographers of American science and culture to students and general readers with an interest in the history of the life sciences, mathematical biology and the social construction of science.

Twenty years ago, researchers wishing to identify contaminated areas in aquatic environments generally took water samples, and analysed them badly (as we have since discovered) for a few "pollutants" which were of topical note at the time (and which could be quantified by the methods then available). Today, the use of aquatic organisms as biomonitors in preference to water analysis has become commonplace, and many national and interna tional programmes exist around the world involving such studies. We believe that this trend will continue, and have complete faith in the methodology (when it is employed correctly). We hope that the following text assists in some part in attaining this goal, such that the quality of our most basic global resource -water - is adequately protected in the future. DAVE PHILLIPS, PHIL RAINBOW England, March 1992 vii Acknowledgements Our thanks for contributions to this book are due to several individuals and groups, for varying reasons. Firstly, a co-authored book is always a triumph, and we trust that the following text is an acceptable compromise of the views of two individual authors, on a complex and developing topic. Secondly, many of the ideas herein have crystallised over the last two decades as the field has grown, and we are individually and collectively grateful to a number of researchers for their insight and assistance."

What sight is more beautiful than a high-energy beach facing lines of rolling white breakers? What battleground is more ferocious than where waves and sand meet? What environment could be more exciting to study than this sandy interface between sea and land? And yet how much do we know about sandy beaches? Open sandy beaches are amongst the most neglected fields of scientific study in the coastal environment. This situation exists despite their great extent along most temperate and tropical coastlines and their value as recreational areas and buffer zones against the sea. The traditional oceanographer does not venture into the surf zone while the terrestrial ecologist stops short at the high water mark. Only a few coastal engineers have grappled with the problem of sand and sediment movement as it influences their construction of harbours and pipelines. The marine biologist on the other hand has regarded estuaries, coral reefs and rocky shores, obviously teeming with life, as more fruitful areas for study than the apparently poor animal life on sandy beaches. Sandy beaches have therefore tended to become a scientific no man's land. Over the last decade this situation has begun to improve. Recent work on high-energy beaches has revealed that they may in fact be rich and productive and fertile areas for study. It has even been suggested that beaches and their adjacent surf zones may constitute viable marine ecosystems.

The Ebro is a typical Mediterranean river characterized by seasonal low flows and extreme flush effects, with important agricultural and industrial activity that has caused heavy contamination problems. This volume deals with soil-sediment-groundwater related issues in the Ebro river basin and summarizes the results generated within the European Union-funded project "AquaTerra." The following topics are highlighted: Hydrology and sediment transport and their alterations due to climate change, aquatic and riparian biodiversity in the Ebro watershed, occurrence and distribution of a wide range of priority and emerging contaminants, effects of chemical pollution on biota and integration of climate change scenarios with several aspects of the Ebro s hydrology and potential impacts of climate change on pollution.

The primary objective of the book is to lay the foundation for a better understanding of the behavior of environmental pollutants and their fluxes with respect to climate and land use changes."

Keith Culver and David Castle Introduction Aquaculture is at the leading edge of a surprisingly polarized debate about the way we produce our food. According to the United Nations Food and Agriculture Organization, aquaculture production has increased 8. 8% per year since 1970, far surpassing productivity gains in terrestrial meat production at 2. 8% in the same period (FAO 2007). Like the 'green revolution' before it, the 'blue revolution' in aquaculture promises rapidly increased productivity through technology-driven - tensi?cation of aquaculture animal and plant production (Costa-Pierce 2002; The Economist 2003). Proponents of further aquaculture development emphasize aq- culture's ancient origins and potential to contribute to global food security d- ing an unprecedented collapse in global ?sheries (World Fish Center; Meyers and Worm 2003; Worm et al. 2006). For them, technology-driven intensi?cation is an - dinary and unremarkable extension of past practice. Opponents counter with images of marine and freshwater environments devastated by intensive aquaculture pr- tices producing unsustainable and unhealthy food products. They view the promised revolutionasascam, nothingmorethanclever marketingbypro?t-hungry ?shfa- ers looking for ways to distract the public from the real harms done by aquaculture. The stark contrast between proponents and opponents of modern aquaculture recalls decades of disputes about intensive terrestrial plant and animal agriculture, disputes whose vigor shows that the debate is about much more than food production (Ruse and Castle 2002).

This volume is derived from a symposium on High Latitude Limnology held during the 23rd Congress of the Societas Internationalis Limnologiae in Hamilton, New Zealand. The symposium stemmed from our belief that an exchange of views between limnologists working in the north and south polar zones would be timely and productive. Over the last decade there has been a major increase in the limnological research effort in Antarctica with the expansion of science programmes there by many nations from both the northern and southern hemispheres. Freshwater research has also continued to develop in the Arctic, stimulated by the need for basic information to assess environmental impacts of the oil industry, mining, urbanisation and other human activities. By bringing together aquatic investigators from both poles we hope to draw attention to the distinctive features that high latitude systems hold in common, and to the marked contrasts between and within each zone. The dominant impression from the assemblage of papers presented here is one of great limnological diversity. The studies include clear, turbid and brown water rivers in the sub Arctic (LaPerriere, Van Nieuwenhuyse & Anderson); chlorophyte dominated streams in the maritime Antarctic (Hawes); streams on the antarctic continent lined with thick cyanobacterial mats (Howard-Williams & Vincent); meromictic waters in the Arctic (Ouellet, Dickman, Bisson & Page) and Antarctic (e. g.

The first edition of the widely praised Marine Protected Areas for Whales, Dolphins and Porpoises, published in 2005, led to numerous new marine protected area proposals and a number of notable conservation successes around the world.

In this completely revised and expanded second edition, new developments in the Mediterranean, Caribbean and Pacific are described, as well as future directions for High Seas protection. New sections show how to design and manage Marine Protected Areas (MPA's) in an ever noisier ocean subject to climate change, increased shipping and hydrocarbon exploration. The process of protected area creation for cetaceans has been accelerated and more than 200 exciting new places are detailed in this edition. This book provides a route map for MPA managers, as well as countries, to meet the ambitious targets for highly protected MPA networks by 2012 and 2020.

Marine Protected Areas for Whales, Dolphins and Porpoises is a key conservation tool and a springboard for worldwide change in human attitudes toward the world ocean where all life originated and where the majority of life on Earth still lives.

'A lake is the landscape's most beautiful and expressive feature. It is earth's eye; looking into which the beholder measures the depth of his own nature. The fluviatile trees next the shore are the slender eyelashes which fringe it, and the wooded hills and cliffs around are its overhanging brows.' 'A field of water betrays the spirit that is in the air. It is continually receiving new life and motion from above.' 'Nothing so fair, so pure, and at the same time so large, as a lake, perchance, lies on the surface ofthe earth. Sky water. It needs no fence. Nations come and go without defiling it. It is a mirror which no stone can crack, whose quicksilver will never wear off, whose gilding Nature continually repairs; no storms, no dust, can dim its surface ever fresh; - a mirror in which all impurity presented to its sinks, swept and dusted by the sun's hazy brush, - this the light dust-cloth, - which retains no breath that is breathed on it, but sends its own to float as clouds high above its surface, and be reflected in its bosom still.' (Henry David Thoreau: 'Walden; or Life in the Woods'. Fourth printing, April 1953, Ri!lehart & Company, New York).

Cell signalling lies at the heart of many biological processes and currently is the focus of intense research interest. In multicellular organisms, it is central to how different types of cell communicate with each other and how they detect and respond to extracellular signals. Intercellular communication is vital to single-celled organisms as well, allowing them to respond to environmental cues and signals.

To date, much of the understanding of signalling mechanisms has come from research on specific cell types (eg mouse lymphocyte and cardiomyocyte) or on organisms in which communication systems such as nervous and endocrine systems are well established. This volume therefore aims to 'fill the gap' by concentrating on 'simple organisms' where the elements of those signalling systems first evolved. Many of the groups covered contain important pathogens or parasites, and the potential for manipulating signalling pathways for therapeutic intervention will be highlighted.

Concentrations of dissolved organic matter (DOM) in lakes are often an order of magnitude greater than concentrations of particulate organic matter; nevertheless, the biogeochemical analysis of DOM is described in only a few textbooks on limnology (most thoroughly by Wetzel). The orgins of dissolved organic substances are largely photosynthetic; DOM is either autochthonously synthesized by littoral and pelagic flora through secretions and autolysis of cellular contents, or allochthonously generated in terrestrial systems of the drainage basin, composing largely of humic substances refractory to rapid microbial degradation. The role of DOM in lacustrine ecosystems, as energy source and system regulator, however, is still poorly known. The aim of this book is: (1) to present state-of-the-art reviews of the role of dissolved autochthonous and allochthonous organic matter in pelagial and littoral zones; and (2) to focus attention on poorly understood but critical topics and hence to provide direction for future research activity.

The Netherlands, like the other industrialized countries in the northern hemisphere, is faced with increas ing environmental pro blems. Being a country rich in water - at least in terms of surface area - and low-lying, problems related to water are as old as the nation itself. The problems in regard to water quality are becoming more acute and complicated due to the progressive rise in air and ground pollution. Glaring examples are the acid rains and contamination of ground-water resources by toxic, and often illegal, waste dumps. Being a delta country, the Netherlands is also very vulnerable to pollution from the*water sources outside the country's national boundaries, i.e. the rivers Rhine, Meuse and ScheIdt. Not only the national but also the international authorities are active in developing measures to both prevent and combat water pollution, and to restore as far as possible the natural balance or a 'reasonable' water quality. The pollution abatement programmes are based on ecological theories; accepting waterbodies as aquatic ecosystems, the authorities concerned try to influence and lead the progresses in the systems to reach or maintain the desired ecosystem structure. Suter II (1981) mentioned a number of reasons why the attempts to use ecosystem theories to assess the impacts on the environment have been disappointing to date. It is clear that one of the reasons for the failure is lack of proper knowledge.

Estuaries exist along the edge of the oceans and seas, and are char acterized by the dilution of sea water by inflowing fresher waters. The motion and interaction of these two types of water (fresh and salt water) determine the salinity distribution within the estuary and that, in turn, affects the organisms residing there. The purpose of this vol ume is to review the status of our understanding of estuarine circu lation and how the circulation patterns affect living and nonliving resources in estuaries. For many years, the primary paradigm for estuarine circulation was the two-layered net or nontidal gravitational circulation pattern first proposed by Dr. Donald Pritchard in his studies of the James River estuary. During the last decade or so, research has focused on the many variations about this theme and the factors that control the transport processes. Many of these aspects are covered in the initial papers in this volume. Water movement, of course, is of interest be cause it transports marine organisms, sediments, and pollutants. Es tuarine circulation has a significant effect on estuarine food chains, and on the distribution and abundance of organisms, such as the American oyster, that are freely transported by the currents during larval stages. The intent is to bring together many of these topics in a single volume. This volume is dedicated to Dr. Donald W."

P. G. Sly Canada Centre Inland Waters, P.O. Box 5050, Burlington, Ontario L7R 4A6, Canada Keywords: symposium, sediments, freshwater Following the highly successful First Symposium work of the symposium: on Research into the Interactions between Sedi- I. Summary of sediment dynamics and distribu- ments and Freshwater held in Amsterdam, in 1976, tion in fluvial and lacustrine environments. it was decided that a second symposium in this 2. Land use effects on sediment yields,and quality. series should be held to continue the discussions on 3. Sediments in mass balance models, including progress in this rapidly expanding field of aquatic total loadings. science. The Second Symposium was held at 4. Sediments as concentrating mechanisms of nat- Queen's University, Kingston, Ontario, between 15 ural and anthropogenic, organic and inorganic and 18 June 1981. It was originally intended to hold materials. the meeting in Toronto a year earlier, but because 5. Release mechanisms, and transfer to biosystem, of difficulties in planning and the high cost of ac- from bed and suspended solids. commodation it was necessary to revise the original 6. Short- and long-term dating methods, spatial program and schedule. and temporai consistency, bioturbation and dif- The symposium received generous support from fusion.

This volume contains the Proceedings of the Third International Conference on Copepoda, held at the British Museum (Natural History) in London during August 1987. The central theme of the conference was the biology of marine planktonic copepods, although the scientific programme was extremely varied reflecting the wide range of life styles adopted by copepods. The three invited symposia held during the conference focussed attention on particular topical areas of research within the field of marine plankton, and also provided reviews of chosen aspects of copepod biology. These symposia were highly successful. The papers they contained were both informative and stimulating and they bring to this volume a lasting significance. Each symposium was organised by its chairman; Bruce Frost (University of Washington) decided on the balance of topics, selected the speakers and introduced the session on 'The biology and taxonomy of Calanus', Roger Harris (Marine Biological Association) performed the same vital role for 'Experimental studies: rate processes in field populations of planktonic copepods', and Howard Roe (Institute of Oceanographic Sciences) for 'Oceanic and deep-sea copepods'. The impact of these papers will be much enhanced by the large number of high quality contributed and poster papers on marine plankton and by the invited review of 'Copepod luminescence' by Peter Herring (Institute of Oceano graphic Sciences). The fascinating review of 'Copepod eyes' by Mike Land FRS (University of Sussex) is not published here."

Reflecting increased interest in the field and its relevance in global environmental issues, Oceanography and Marine Biology: An Annual Review, Volume 47 provides authoritative reviews that summarize results of recent research in basic areas of marine research, exploring topics of special and topical importance while adding to new areas as they arise. This volume, part of a series that regards the all marine sciences as a complete unit, features contributions from experts involved in biological, chemical, geological, and physical aspects of marine science. These features along with the inclusion of a full color insert and an extensive reference list, make the text an essential reference for researchers and students in all fields of marine science.

The global scale of alien species invasions is becoming more and more evident in the beginning ofthe new millennium. Though the problem ofbiological invasions became a rapidly growing research area, there are large gaps still, both geographically and the matically, to be filled in the near future. This book is the first attempt to provide an overall picture of aquatic species invasions in Europe. Its geographical scope stretches from Irish waters in the west to Volga River and the Caspian Sea in the east, and from Mediterranean in the south up to the Arctic coast of Europe. Not all parts of the continent could be equally covered, as in some countries species invasions are not studied yet. The book tends to represent the array of all major European aquatic systems on the broadest geographical and ecological scope possible from fully saline seas, semi-enclosed brackish water bodies and coastallagoons to freshwater lakes, major river systems and waterways. The key objectives include the present status and impacts caused by non-native aquatic species in European waters. Please note that lengthy species lists submitted for publication and additional informa tion were put on the Internet, as the electronical version of these tables benefits from computer assisted search for data (http: //www. ku. lt/nemo/EuroAquaInvaders. htm). Altogether more than 100 scientists from 24 countries have joined to synthesize the available information on bioinvasions. However, the book does not claim to be fully comprehensive."