Charles Darwin (1809 1882) first published this work in 1868 in two
volumes. The book began as an expansion of the first two chapters
of On the Origin of Species: 'Variation under Domestication' and
'Variation under Nature' and it developed into one of his largest
works; Darwin referred to it as his 'big book'. In volume 2,
concerned with how species inherit particular characteristics,
Darwin first published his 'provisional hypothesis' of pangenesis.
This theory of 'gemmules' was not met with much acceptance and
today is not valuable as scientific explanation, but it was
important in laying down the key questions that needed to be
answered regarding the processes of genetic inheritance. Darwin
also used volume 2 to challenge the theories of evolution by
design, expounded by the botanist Asa Gray. Darwin's arguments were
some of the very first in a long debate that remains hot today.
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