TheInternationalConferencesonEphemeroptera(Mayflies)andSymposiaonPlecoptera (Stoneflies)areheldeverythreeyears,indifferentpartsoftheworld. Theseeventsallow specialistsfromdifferentcountriestointeractandpresenttheresultsoftheirlatest investigations. TheIXInternationalConferenceonEphemeropteraandXillInternationalSymposiumon Plecoptera,wereheldAugust16-21,1998,andAugust20-23,1998,respectively,inTaffdel Valle,Tucuman,Argentina,withajointsymposiumonAugust20,1998. Theseeventswerehos- tedbythe"InstitutoSuperiordeEntomologfa"FacultaddeCienciasNaturales,fromtheNational UniversityofTucuman,andattendedbyapproximately80specialistsfrom25countries. Atpresent,whenthebiodiversitycrisisandthepressuresonfresh-waterenvironmentsand theirinhabitantsareworsethaneverbefore,theinformation,discussionsandguidelinescoming outofeventsofthiskindarebecomingmoreandmoreimportant. The54papersincludedinthisvolumeareamongthosepresentedduringthemeetings, andacceptedafterpeerreviewbyinternationalspecialists. Thepapersaregroupedinfivelooselydefinedsections(exceptthefirstthatcorresponds toapaneldiscussion),althoughseveralofthepaperscouldexceedthesubjectboundaries wheretheyarelocated. Iwouldliketothankthemanyindividualsandinstitutionsthathelpedwithboththis bookandtheorganizationoftheevents,namely: Themembersoftheorganizingcommittee:H. R. Fernandez,M. G. Cuezzo, F. Romero, C. Molineriand C. Nieto. AlsocollaboratingwereM. Ceraolo,1. Chocobar,M. Guzmande Tome,S. Moro,M. Orce,V. Manzoandmanyvolunteersandstudentstoonumeroustodetail here. Withouttheiruntiringefforts,theeventssimplywouldnothavebeenpossible. Thefollowingpersonsactedasmanuscriptreviewers:J. Alba-Tercedor,J. V. Arnekleiv, R. Baumann,J. E. Brittain, 1. C. Campbell,J. M. Elouard,J. F. Flannagan,R. W. Flowers, C. Froehlich,E. Gaino,M. T. Gillies,P. Goloboff,P. M. Grant,M. D. Hubbard, Y. Isobe,J. Jackson,N. N. Kapoor,P. Landolt, W. P. McCafferty, 1. McLellan,R. Nelson, W. L. Peters,G. Pritchard,M. Sartori,!. Sivec,1. Stanford, K. W. Stewart,D. Studemann,B. Stark,P. Suter,B. Sweeney,S. Szczytko,andP. Zwick. Theseinstitutionsprovidedtheirinstitutionaland/oreconomicsupport:Facultadde CienciasNaturaleseInstitutoMiguelLillo,UniversidadNacionaldeTucuman;ConsejoNa- cionaldeInvestigacionesCientificasyTecnicas(CONICET);ConsejodeInvestigaciones, UniversidadNacionaldeTucuman(CIUNT);Fundaci6nAntorchas,Fundaci6nMiguelLillo; Direcci6ndeTurismo,MunicipalidaddeTaffdelValle. vii viii Preface ThePermanentCommitteeonEphemeropteraConferencesprovidedpartialfinancial supportforfellowshipsforstudentsfromtheCzechRepublic,ChinaandSlovenia;andthe NorthAmericanBenthologicalSocietyforstudentsfromBoliviaandVenezuela. Thecompletionofthisbookwouldnothavebeenpossiblewithouttheinvaluableeffort ofGustavoSanchez,whomadethedigitalworkoftheoriginalsandthecoverdesign, C. Molineriand C. Nietowhopatientlyreviewedthefinalcopies,andvaluableadvicefrom MaryAnnMcCarraandRobertWheeler. ThecoverillustrationsweredonebyS. RoigJu- nen!(Ephemeroptera)and A. Dupuy(Plecoptera). EduardoDominguez ConvenorandEditor August,2000 CONTENTS STATUS OF THE KNOWLEDGE OF EPHEMEROPTERA IN THE WORLD IntroductiontothePanelDiscussion . 1. Alba-Tercedor StatusoftheKnowledgeofEphemeropterainNortheastAsiaandGuidelinesfor FutureResearch...3 y. 1. Bae TheCurrentStatusofEphemeropteraBiologyinAustralia...7 I. C. Campbell KnowledgeoftheAfrican-MalagasyMayflies ...13 J. M. Elouard TheGentleQuest: 200YearsinSearchofNorthAmericanMayflies...21 W. P. McCafferty TheStatusoftheTaxonomyoftheMayfly(Ephemeroptera)FaunaofSouthAmerica. . 37 M. L. Pescador,M. D. Hubbard,andM. delC. Zuniga TheEphemeropteraofNewZealandandNewCaledonia...43 W. L. Peters CurrentKnowledgeofMayflyResearchinEurope(Ephemeroptera)...47 M. Sartori StatusoftheSystematicKnowledgeandPrioritiesinEphemeropteraStudies: theOrientalRegion '...53 T. Soldan ECOLOGYANDBEHA~OUR LifeCycleandAnnualProductionof Caenissp(Ephemeroptera,Caenidae) inLakeEscondido(Bariloche,Argentina)...67 D. A. AnonSuarezandR. 1. Albarino ix x Contents EffectsofRotenoneTreatmentonMayflyDriftandStandingStocks inTwoNorwegianRivers...77 1. V. Arnekleiv,D. Dolmen,and L. Rj(jnning LongitudinalDistributionoftheMayfly(Ephemeroptera)Communities attheChocancharavaRiverBasin(Cordoba,Argentina)...89 M. delC. Corigliano,C. M. Gualdoni, A. M. Oberto,andG. B. Raffaini EmergenceofEphemeropterafromtheAssiniboineRiver,Canada...97 1. F. Flannagan,1. Alba-Tercedor,R. G. Lowen,andD. G. Cobb MayfliesasFood 107 P. M. Grant SeasonalVariationofEphemeropterainFourStreamsofGuatopoNational Park,Venezuela...:...125 V. Maldonado,B. Perez,andC. Cressa AbundanceandAltitudinalDistributionofEphemeropterainan Andean-PatagoneanRiverSystem(Argentina)...

Japanese Encephalitis (JE), a mosquito borne disease, is the leading cause of viral encephalitis in 14 Asian countries due to its epidemic potential, high case fatality rate and increased possibility of lifelong disability in patients who recover from this dreadful disease. In spite of seriousness of the disease, still only few books are available for ready reference. Hence, this book will be useful for students, entomologists, paramedical staff and vector control managers in public health. The objective of the book is to disseminate the knowledge gained by the author from ecological studies on JE vectors undertaken in 2 endemic and 2 non endemic areas of Northern India over a period of last 15 years (1998-2012) of research in the field of ecology of mosquito vectors of JE virus initially at National Institute of Communicable Diseases (Ministry of Health & Family Welfare, Government of India), Delhi and later at Jamia Millia Islamia, a Central University, Delhi, to anyone who wishes to curtail death of children due to this dreaded disease. Of the thousand suspected JE deaths in India annually, more than 75% is contributed by Northern India wherein disease transmission failed to be explained based on entomological evidence due to inadequate mosquito surveillance tool used in determining JE vector density. In order to overcome the above problem, Dr Bina Pani Das, the author of this book, developed "BPD hop cage method", a simple, cost effective, and operationally feasible surveillance tool specially designed to capture predominantly day resting adult Cx. tritaeniorhynchus mosquitoes, the principal JE vector species in the country from land and aquatic vegetation.

Cyanobacteria have existed for 3.5 billion years, yet they are still the most important photosynthetic organisms on the planet for cycling carbon and nitrogen. The ecosystems where they have key roles range from the warmer oceans to many Antarctic sites. They also include dense nuisance growths in nutrient-rich lakes and nitrogen-fixers which aid the fertility of rice-fields and many soils, especially the biological soil crusts of arid regions. Molecular biology has in recent years provided major advances in our understanding of cyanobacterial ecology. Perhaps for more than any other group of organisms, it is possible to see how the ecology, physiology, biochemistry, ultrastructure and molecular biology interact. This all helps to deal with practical problems such as the control of nuisance blooms and the use of cyanobacterial inocula to manage semi-desert soils. Large-scale culture of several organisms, especially "Spirulina" ("Arthrospira"), for health food and specialist products is increasingly being expanded for a much wider range of uses. In view of their probable contribution to past oil deposits, much attention is currently focused on their potential as a source of biofuel.

Please visit http: //extras.springer.com/to view Extra Materials belonging to this volume.

Thisbook complements thehighly successful "Ecology of Cyanobacteria" and integrates the discoveries of the past twelve years with the older literature.

"

British mosquitoes are often overlooked by entomologists in favour of their wilder, tropical cousins. This book brings together all of the current research and information on British mosquitoes, providing a comprehensive, accessible guide to the study and identification of British species. Chapters cover life histories, identification and habitat, accompanied by detailed illustrations. Detailed keys for the identification of eggs, larvae, pupae and adults form the centre of the book, which also includes practical guidance for studying mosquitoes, including where to find them and how to recognise them in all stages of their life cycle. This book is a valuable resource for anyone seeking to broaden their knowledge of the British mosquito, from those with an amateur interest, to students and professionals seeking to publish research on the species. This is a reprint of the first edition published in 1990 (ISBN 0-85546-275-2).

An unprecedented look at the complex and beautiful world of underground ant architecture Walter Tschinkel has spent much of his career investigating the hidden subterranean realm of ant nests. This wonderfully illustrated book takes you inside an unseen world where thousands of ants build intricate homes in the soil beneath our feet. Tschinkel describes the ingenious methods he has devised to study ant nests, showing how he fills a nest with plaster, molten metal, or wax and painstakingly excavates the cast. He guides you through living ant nests chamber by chamber, revealing how nests are created and how colonies function. How does nest architecture vary across species? Do ants have "architectural plans"? How do nests affect our environment? As he delves into these and other questions, Tschinkel provides a one-of-a-kind natural history of the planet's most successful creatures and a compelling firsthand account of a life of scientific discovery. Offering a unique look at how simple methods can lead to pioneering science, Ant Architecture addresses the unsolved mysteries of underground ant nests while charting new directions for tomorrow's research, and reflects on the role of beauty in nature and the joys of shoestring science.

Biocontrol is among the most promising methods for a safe, environmentally benign and sustainable pest control. Microbial pesticides offer a great potential, and it is anticipated that they will become a substantial part of the use of all crop protection products. Their development and commercialization, however, has been difficult and with many failures. In this book a rational and structured roadmap has been designed for the development and commercialization of microbial pest control products for the control of arthropod pests. The building blocks of the entire process are identified and essential aspects highlighted. Biopesticides based on entomopathogenic bacteria, fungi, viruses and nematodes are elaborately discussed. This systematic roadmap with a strong focus on economics and market introduction will assist academic researchers and industrial developers of biopesticides in accomplishing their goal: the development of successful cost-effective microbial pesticides.

Throughout the 1950s, methods of experimental analysis led to a great increase in knowledge of the machinery of insect flight. In this 1957 monograph, Dr Pringle brings together the results of his investigations of the mechanics of wing motion, the structure and physiology of flight muscle, aerodynamics, sense organs and nervous co-ordination, and shows how a common structural plan has been perfected during the course of evolution to produce the variety of form and function found in the different insect orders. Of particular interest is the special type of muscle found only in the higher insects and responsible for the very high wing-beat frequencies of some flies and bees. Many different lines of approach have contributed to the understanding of insect flight, and the subject is of great interest to entomologists, comparative physiologists and biological engineers.

Low temperature is a major environmental constraint impacting the geographic distribution and seasonal activity patterns of insects. Written for academic researchers in environmental physiology and entomology, this book explores the physiological and molecular mechanisms that enable insects to cope with a cold environment and places these findings into an evolutionary and ecological context. An introductory chapter provides a primer on insect cold tolerance and subsequent chapters in the first section discuss the organismal, cellular and molecular responses that allow insects to survive in the cold despite their, at best, limited ability to regulate their own body temperature. The second section, highlighting the evolutionary and macrophysiological responses to low temperature, is especially relevant for understanding the impact of global climate change on insect systems. A final section translates the knowledge gained from the rest of the book into practical applications including cryopreservation and the augmentation of pest management strategies.

The first section of the book includes information about how tourism-related infrastructure and activities promote biological invasions, including key pathways for non-native invasive species introductions. This section provides case studies of different organisms that are known to be introduced and/or promoted by tourism in different ecosystems or regions. The second section elaborates on known and potential impacts of invasive species on tourism and recreation, including how they may affect, positively or negatively, the economic revenue from tourism, tourist access, recreation, aesthetic values and tourists' perceptions. The last section focuses on management and policy, covering aspects of how visitors perceive invasive species and their willingness to manage them, biosecurity measures to prevent invasion related to tourism, as well as potential policy options moving forward. The book draws on a number of examples across multiple taxa, landscapes and regions of the world.

In 1974 when I published my book, Biological Mechanism of Attachment, not many pages were required to report on the attachment devices of insect cuticles. As in most fields of research, our knowledge on this specific subject has simply exploded. Dr. Stanislav N. Gorb now describes the present day level of our knowledge, to which he has personally contributed so much, and a research team working on biological microtribology has gradually developed, also. With modern methods of measurement it is possible to enter the structure - function relationship much more deeply, even down to a molecular level, which was not possible two and a half decades ago. It is a well known fact that, in biology, the more sophisticated the measuring method, the greater the achievement of biological fundamental research, and its resulting evidence. Our knowledge remains at a certain level until new methods once more permit a forward leap. Biological knowledge develops in the form of a stepped curve rather than linear, as reflected in the studies carried out on the attachment devices of insect cuticles.

Egg Parasitoids in Agroecosystems with emphasis on Trichogramma was conceived to help in the promotion of biological control through egg parasitoids by providing both basic and applied information. The book has a series of chapters dedicated to the understanding of egg parasitoid taxonomy, development, nutrition and reproduction, host recognition and utilization, and their distribution and host associations. There are also several chapters focusing on the mass production and commercialization of egg parasitoids for biological control, addressing important issues such as parasitoid quality control, the risk assessment of egg parasitoids to non-target species, the use of egg parasitoids in integrated pest management programs and the impact of GMO on these natural enemies. Chapters provide an in depth analysis of the literature available, are richly illustrated, and propose future trends.

Insect Learning is a comprehensive review of a new field. Until recently, insects were viewed as rigidly programmed automatons; now, however, it is recognized that they can learn and that their behavior is plastic. This fundamental change in viewpoint is causing a re-examination of all aspects of the relationship between insects and their environment. This change in perspective is occurring at a time of heightened interest in brain function in both vertebrates and invertebrates. Insects potentially play a major role in this expanding area. Because of their experimental tractability and genetic diversity, they provide unique opportunities for testing hypotheses on the ecology and evolution of learning. As organisms of economic importance, they are perennial objects of research by both basic and applied scientists. Insect Learning covers both social and non-social insects from multiple perspectives. The book covers mechanisms; syntheses of work on physiology, behavior, and ecology; and micro- and macroevolution. The concluding section discusses future directions for research, including applications to pest management.

Due to the attractiveness of butterflies, and their usefulness as model systems for biological questions, there has been a considerable amount of material written on butterfly biology, largely in Europe. This book synthesizes all relevant and recent knowledge in the field, which is a must for those making use of this taxonomic group as a model system. It is divided into five major parts which deal with habitat use, population ecology and genetics, evolutionary ecology, distribution and phylogeny, and global change and conservation. There are growing numbers of scientific projects and networks in Europe in which the use of butterflies as tools and targets for conservation is central, and application of knowledge is closely related to European cultural landscapes. However, the chapters can also be applied to a wide geographic scope. Written by an international team of experts, this timely book is suitable for students, researchers and enthusiasts.

The subject area embraced by the term "biological control" in its classical sense is very broad indeed. The term itself was apparently first used in 1919 by the late Harry S. Smith, and was then used specifically in reference to the suppression of insect populations by the actions of their indigenous or introduced natural enemies. The California school of biological control specialists who followed in Smith's footsteps have traditionally differentiated "natural" biological control (by indigenous natural enemies) and "applied" biological control (by man-introduced natural enemies). Subsequently, the philosophy broadened beyond the original narrow concern with population suppression of insects (and especially pest insects), to embrace directed activities against mites or other arthropod pests, various invertebrate and vertebrate pests, weeds, and organisms producing disease in humans or their domestic animals and plants. The techniques used in these activities also multiplied beyond the original concern with natural enemies. The subjects area discussed in this book is, at the same time, broader and more restricted than that covered in other books on "biological control. " On the one hand, the treatment here is restrictive in that, with rare exception, we have limited ourselves to dealing only with ideas and examples involving the suppression of insect pests through human activity or intervention in the environment.

Most neurobiological research is performed on vertebrates, and it is only natural that most texts describing neuroanatomical methods refer almost exclusively to this Phylum. Nevertheless, in recent years insects have been studied intensively and are becoming even more popular in some areas of research. They have advantages over vertebrates with respect to studying genetics of neuronal development and with respect to studying many aspects of integration by uniquely identifiable nerve cells. Insect central nervous system is characterized by its compactness and the rather large number of nerve cells in a structure so small. But despite their size, parts of the insect eNS bear structural comparisons with parts of vertebrate eNS. This applies particularly to the organization of the thoracic ganglia (and spinal cord), to the insect and vertebrate visual sys tems and, possibly, to parts of the olfactory neuropils. The neurons that make up these areas in insects are often large enough to be impaled by microelectrodes and can be injected with dyes. Added to advantages of using a small eNS, into which the sensory periphery is precisely mapped, are the many aspects of insect behaviour whose components can be quan titized and which may find both structural and functional correlates within clearly defined regions of neuropil. Together, these various features make the insect eNS a rewarding object for study. This volume is the first of two that describe both classic and recent methods for neuroanatomical research on insect eNS."

Mention the words 'arthropod cuticle' to most biologists and they usually provoke a glazed expression. This is because the cuticle is commonly regarded as an inert substance. It is hoped that this book will dispel this fallacy. The study of cuticle in its proper context now involves many of the wider aspects of biology which are currently in vogue (e. g. how a hormone like ecdyson induces a specific enzyme like dopa decarboxylase; the unsolved major problem of cell gradient and polarity; the involvement of cyclic AMP in hormonal mechanisms; the extra cellular control of cuticular enzymes, of the mechanical proper ties of cuticle structural proteins, and of the orientation of fibrous molecules; and the relation of chromosome puffing to the synthesis of specific proteins). Studies on cuticle demand a variety of techniques, and examples of the following are illustrated in this book (fluorescence, phase contrast, polariza tion and Nomarski interference microscopy; infrared absorp tion; transmission and scanning electron microscopy; autora diography analyzed by electron microscopy; negative staining in the electron microscope; optical diffraction, high angle X-ray diffraction, low angle X -ray diffraction and selected area electron diffraction). I am well aware that the biophysical parts of this book are less incomplete than other aspects. A developmental biologist or a biochemist would have further elaborated other parts ofthe subject matter. Only one previous author, RICHARDS (1951)hasdevoted a book to arthropod cuticle."

The literature is still one of our biggest frustrations to-day. There is, in one sense, too much of it, and in another not enough - for there are insufficient and inadequate published guidelines through this jungle. Last year two excellent books for students of ecological chemistry were published, one in France and one in England. The concordance of the references was a mere overall 3% rising to 7% in the chapters on pheromones. Even in the computer age, the channel remains a formidable barrier to the rapid exchange of biological information. At the present time we are in urgent need of compilations similar to John Feltwell's "The Large White Butterfly"; since the literature has become virtually unmanageable. This insect is now a demonstration object in the sixth form schoolroom; an experimental "rabbit" in the University laboratory; a test animal in virus and bacterial research projects; a tool for the study of flight mechanisms, migration, plant biochemistry, hormones, genetics, allergies, pigments, mimicry, etc. , etc. John Feltwell has, by this massive compilation, rendered us a great service - in fact he has given us a present of 4,000 hours of library time spent in 50 different libraries in seven countries. In the process he has collected 8000 references to the Large White. Of these, 4000 have been selected, and we are given a brief indication of their contents.

The fifth and last Volume of this IPMD series reviews, in a multi-disciplinary approach, recent achievements in crop protection and integrated management of arthropod pests. The volume is organized in a first Section covering IPM in citrus productions, a Second one dealing with advacements in the integratioon of management technologies and a last Section covering mites and their biological control agents. As for the previous volumes, we attempted to provide an informative coverage for a broad range of agricultural systems and situations. The chapters are mainly organized and centered on crops, with a particular emphasis on citrus. This is one of the main crops in which biological control and IPM approaches were tested successfully, and the experience gained herein may indeed result helpful for IMP efforts deployed worldwide on other crops and/or cropping systems. Chapters in Section 2 review the integration of insect and disease management options in pecan crops, the application of remote sensing technologies, the status of knowledge about plant defense compounds and their potentials. For IPM of invasive species, an update is provided on the experience gained on the Red Palm Weevil (RPW) in Egypt. Long-term technological solutions for IPM in forests and park areas through aerial treatments with Bt spores concludes this Section. Finally, in Section 3, updated reviews about biological control agents of mites are provided.

The function of insect blood cells has long puzzled scientists. This much-needed volume, first published in 1979, attempts to explain the forms and functions of insect hemocytes, creating a solid basis from which work in the field may proceed. Insect Hemocytes had its origins in a symposium held at the XV International Congress of Entomology (1976). It is a coherent presentation whose contributors were some of the most eminent specialists in the field.

This volume is an outgrowth of a Symposium entitled "Evolution of Escape in Space and Time" held at the XV International Congress of Entomology in Washington, D. C., USA in August, 1976. The choice of topic was prompted by recent advances in evolutionary ecology and the apparent suitability of insect migration and dia pause as appropriate material for evolutionary studies. In the event, that choice seems amply justified as I hope a perusal of these papers will show. These Sympos ium papers hardly cover the topic of the evolution of escape mechanisms exhaustively, and I am sure everyone will have his favorite lacuna. Some of the more obvious ones are indicated by Professor Southwood in his Concluding Remarks at the end of the book. The purpose of the Symposium, however, was not complete coverage, but rather to indicate the potential inherent in insect migration and diapause for the study of evolutionary problems. In that I think we have succeeded reasonably well. These papers are expanded and in some cases somewhat altered versions of the papers delivered in Washington. This has allowed greater coverage of the topics in question. I suggested a format of a general overview of a topic emphasizing the author's own research con tributions. In general the papers follow this outline although emphases vary. Two of the authors, Dr. Rainey and Dr. Lumme, were unable to attend the Symposium. Dr. Rainey's paper was read by Mr. Frank Walsh, but Dr."

Social insects are amongst the most successful species. This study concentrates on the production ecology of ants and termites. Ants and termites are highly socialised and their groupings in their most developed form enable them to function as large organisms comparable with the larger mammals in their influence in ecosystems. In spite of their social mode of life, quantitative investigations are not easy and this volume emphasises the importance of developing reliable techniques. The account deals critically with production, food and feeding habits, respiration and energy flow, nutrient dynamics and the role of ants and termites in ecosystems. The body of information is not only an invaluable summary and synthesis of work already done, but presents a concise statement of what now needs to be done and what means should be employed. The volume is for social insect ecologists and entomologists and for all who teach ecology, because it contains valuable data and ideas for the general ecologist.

Insects are the most diverse and abundant animals that share our world, and conservation initiatives are increasingly needed and being implemented globally, to safe guard the wealth of individual species. This book provides sufficient background information, illustrated by examples from many parts of the world, to enable more confident and efficient progress towards the conservation of these ecologically indispensable animals. Writing for graduate students, academic researchers and professionals, Tim New describes the major ingredients for insect species management and conservation, and how these may be integrated into effective practical management and recovery plans.

In his preface to early editions of this book, the late Dr. A. D. Imms said that he intended it to be an elementary account of entomology as a branch of general biology. He had especially in mind the needs of university students of zoology and agriculture, as well as those intending later to specialize in entomology, and he suggested that the book might also interest teachers of advanced biology'in schools. These general aims and the balance between the different aspects of the subject have changed little in this and in our previous revision. We have, however, tried to bring the present edition up to date on the lines of our revised tenth edition of Imms' General Textbook of Entomology, published in 1977. The text has been entirely re-set and eleven illustrations have been replaced by new figures. The same orders . of insects are recognized as in the last edition, but the sequence in which the Endopterygote groups appear has been changed to reflect more accurately their probable evolutionary relationships. Many small changes and some addi tions have been made in the physiological sections, the chapter on the origin and phylogeny of insects has been rewritten, and a new bibliography provides a selection of modern references for the in tending specialist. It has been our object to make these alterations without materially increasing the length of the book or its level of difficulty."

For nearly 50 years, pest control was mostly based on broad-spectrum conv- tional insecticides such as organochlorines, organophosphates, carbamates and pyrethroids. However, the severe adverse effects of pesticides on the environment, problems of resistance reaching crisis proportions and public protests led to stricter regulations and legislation aimed at reducing their use. Ways to reduce the use of synthetic pesticides in plant protection and to use more alternative and novel me- ods for pest control or biorational control are the challenges of pest control for the twenty-first century. The term biorational (biological + rational) pesticides can be defined as the use of specific and selective chemicals, often with a unique modes of action, that are compatible with natural enemies and the environment, with minimal effect on n- target organisms. Biorational control is based on a diversity of chemical, biological and physical approaches for controlling insect pests which results in minimum risk to man and the environment.