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Root Hairs - Cell and Molecular Biology (Hardcover, 2000 ed.)
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Root Hairs - Cell and Molecular Biology (Hardcover, 2000 ed.)
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Root hairs are tip-growing cells that originate from epidennal
cells called trichoblasts. Their role may be simply thought of as
extending the surface area of the root to facilitate absorption of
nutrients and water. However, as you will see in this book, the
root hair is far more than that. To an increasingly larger number
of plant biologists, the root hair is a model cell. It grows in
much the same way as a pollen tube, by sending vast numbers of
vesicles containing cell wall precursors to a rounded apical dome,
the tip. Once the trichoblast becomes committed to root hair
fonnation, it no longer divides. The root hair cell has a migrating
nucleus and a complex cytoskeleton. It has a varied cell wall. It
is easy to observe through differential interference contrast
microscopy because there are no other cells around it to disturb
the image. Cytoplasmic streaming is exceptionally clear, and
amyloplasts and even mitochondria and endoplasmic reticulum can be
seen without reporter labelling in some species. Root hair mutants
are easy to distinguish and catalogue. Plant honnones are involved
in their growth and development. It is thus an almost ideal plant
cell for experimental manipulation and observation. The root hair
is also involved in interactions with soil microbes, as you will
learn from later chapters of the book.
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