Now with over 90% new citations, the text emphasises the general principles that apply broadly across taxa and modalities. The topics covered range from the physics and physiology of signal production, propagation, and reception, to complications arising when sender and receiver do not have identical interests during communication.

What cognitive abilities underpin the use of tools, and how are tools and their properties represented or understood by tool-users? Does the study of tool use provide us with a unique or distinctive source of information about the causal cognition of tool-users? Tool use is a topic of major interest to all those interested in animal cognition, because it implies that the animal has knowledge of the relationship between objects and their effects. There are countless examples of animals developing tools to achieve some goal-chimps sharpening sticks to use as spears, bonobos using sticks to fish for termites, and New Caledonian crows developing complex tools to extracts insects from logs. Studies of tool use have been used to examine an exceptionally wide range of aspects of cognition, such as planning, problem-solving and insight, naive physics, social relationship between action and perception. A key debate in recent research on animal cognition concerns the level of cognitive sophistication that is implied by animal tool use, and developmental psychologists have been addressing related questions regarding the processes through which children acquire the ability to use tools. In neuropsychology, patterns of impairments in tool use due to brain damage, and studies of neural changes associated with tool use, have also led to debates about the different types of cognitive abilities that might underpin tool use, and about how tool use may change the way space or the body is represented. Tool Use and Causal Cognition provides a new interdisciplinary perspective on these issues with contributions from leading psychologists studying tool use and philosophers providing new analyses of the nature of causal understanding A ground-breaking volume which covers several disciplines, this volume will be of interest to psychologists, including animal researchers and developmental psychologists as well as philosophers, and neuroscientists.

Do the movements of animals, including humans, follow patterns that can be described quantitatively by simple laws of motion? If so, then why? These questions have attracted the attention of scientists in many disciplines, and stimulated debates ranging from ecological matters to queries such as 'how can there be free will if one follows a law of motion?' This is the first book on this rapidly evolving subject, introducing random searches and foraging in a way that can be understood by readers without a previous background on the subject. It reviews theory as well as experiment, addresses open problems and perspectives, and discusses applications ranging from the colonization of Madagascar by Austronesians to the diffusion of genetically modified crops. The book will interest physicists working in the field of anomalous diffusion and movement ecology as well as ecologists already familiar with the concepts and methods of statistical physics.

The editors utilize their 50 years of combined experience in professional engagement with the behaviour and ecology of wild felids to draw together a unique network of the world's most respected and knowledgeable experts. For the first time, this inter-disciplinary research programme is brought together within a single volume.
Beginning with a complete account of all 36 felid species, there follow 8 comprehensive review chapters that span all the topics most relevant to felid conservation science, including evolution and systematics, felid form and function, genetic applications, behavioural ecology, management of species that come into conflict with people and control of international trade in felid species, conservation tools/techniques, ex situ management, and felid diseases. 19 detailed case studies then delve deeply into syntheses of the very best species investigations worldwide, written by all the leading figures in the field. These chapters portray the unique attributes of the wild felids, describe their fascinating (and conflicting) relationship with humans, and create an unparalleled platform for future research and conservation measures. A final chapter analyses the requirements of, and inter-disciplinary approaches to, practical conservation with cutting-edge examples of conservation science and action that go far beyond the cat family.

The editors utilize their 50 years of combined experience in professional engagement with the behaviour and ecology of wild felids to draw together a unique network of the world's most respected and knowledgeable experts. For the first time, this inter-disciplinary research programme is brought together within a single volume.
Beginning with a complete account of all 36 felid species, there follow 8 comprehensive review chapters that span all the topics most relevant to felid conservation science, including evolution and systematics, felid form and function, genetic applications, behavioural ecology, management of species that come into conflict with people and control of international trade in felid species, conservation tools/techniques, ex situ management, and felid diseases. 19 detailed case studies then delve deeply into syntheses of the very best species investigations worldwide, written by all the leading figures in the field. These chapters portray the unique attributes of the wild felids, describe their fascinating (and conflicting) relationship with humans, and create an unparalleled platform for future research and conservation measures. A final chapter analyses the requirements of, and inter-disciplinary approaches to, practical conservation with cutting-edge examples of conservation science and action that go far beyond the cat family.

Comprising a substantial part of living biomass on earth, ants are integral to the functioning of terrestrial ecosystems. More than 12,500 species have been described to date, and it is estimated that perhaps as many still await classification.
Ant Ecology explores key ecological issues and new developments in myrmecology across a range of scales. The book begins with a global perspective on species diversity in time and space, and examines interactions at the community level before describing the population ecology of these social insects. The final section covers the recent ecological phenomenon of invasive ants: how they move across the globe, invade, affect ecosystems, and are managed by humans. Each chapter links ant ecology to broader ecological principles, provides a succinct summary, and discusses future research directions. Practical aspects of myrmecology, applications of ant ecology, debates, and novel discoveries are highlighted in text boxes throughout the volume. The book concludes with a synthesis of the current state of the field and a look at exciting future research directions. The extensive reference list and full glossary are invaluable for researchers, and those new to the field.

Offers the first comprehensive resource to document the particular needs of a given species in captivity. Each species is dealt with by specialists who are fully familiar with their animals, writing to a consistent template to facilitate easy navigation. Each chapter in Part 2 addresses specific species-typical behaviours associated with pain and distress. The third section contains a series of easy reference ethograms that will aid understanding, and potentially quantification of i.e., baboon, bird, or bovine behavior. The book will be useful not only in laboratory settings, but any situation where an animal is kept in captivity, from veterinary clinics to zoos and sanctuaries. Edited by internationally recognized experts in animal behaviour and behavioural management.

Why do males and females frequently differ so markedly in body size and morphology?
Sex, Size, and Gender Roles is the first book to investigate the genetic, developmental, and physiological basis of sexual size dimorphism found within and among the major taxonomic groups of animals. Carefully edited by a team of world-renowned specialists in the field to ensure a coherence of style and approach between chapters, it presents a compendium of studies into the evolution, adaptive significance, and developmental basis of gender differences in body size and morphology. Adaptive hypotheses allude to gender-specific reproductive roles and associated differences in trophic ecologies, life history strategies, and sexual selection. This "adaptationist" approach is balanced by more mechanistic studies of the genetic, developmental and physiological basis of sexual size dimorphism to provide a comprehensive and authoritative overview of the subject. Throughout the volume the emphasis is on sexual dimorphism in overall size; however, the scope of enquiry encompasses gender differences in body shape, the size and structure of secondary sexual characteristics, patterns of growth (ontogeny), and patterns of gene regulation.
This advanced, research level text is suitable for graduate level students and researchers in the fields of evolutionary biology, behavioral ecology, physiology, developmental biology, and genetics. It will also be of relevance and use to non-biologists from fields such as anthropology and gender studies.
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Communication is essential for all forms of social interaction, from parental care to mate choice and cooperation. This is evident for human societies but less obvious for bacterial biofilms, ant colonies or flocks of birds. The major disciplines of communication research have tried to identify common core principles, but syntheses have been few because historical barriers have limited interaction between different research fields.
Sociobiology of Communication is a timely and novel synthesis. It bridges many of the gaps between proximate and ultimate levels of analysis, between empirical model systems, and between biology and the humanities. The book offers the complementary approaches of a distinguished group of authors spanning a large diversity of research programs, addressing, for example, the genetic basis of bacterial communication, dishonest communication in insect societies, sexual selection and network communication among colonial vertebrates. Other chapters explore the role of communication in genomic conflict and self-organisation, and how linguistics, psychology and philosophy may ultimately contribute to a biological understanding of human mate choice and the evolution of human societies.
This highly interdisciplinary book highlights key examples of modern research to explore the genetic, neurobiological, physiological, chemical and behavioural basis of social communication. It identifies where consensus on the general principles is emerging and where the major future challenges are to be found. The book is therefore suitable for both for graduate students and professionals in evolutionary biology and behavioural ecology seeking novel inspiration, and for a wideracademic audience, including social and medical scientists who would like to explore what evolutionary approaches can offer to their fields.

The Racehorse: A Veterinary Manual has become the definitive text for primary care of the Thoroughbred racehorse. Written by one of the world’s leading racehorse veterinary clinicians, it sets out best practice standards of diagnosis and management of all the major conditions likely to be encountered in racehorse clinical practice, as well as comprehensively reviewing subjects as diverse as pre-purchase assessment and exercise physiology. This second edition has been thoroughly updated and introduces new chapters on a range of topics including injury risk assessment and electrolyte and fluid therapy, expanded sections on nutrition, rehabilitation and sales radiography as well as an array of new images and ready reference charts. The Racehorse: A Veterinary Manual remains an invaluable resource for both clinicians and non-veterinarians in the racing industry.

A short introduction to animal behaviour, investigating a whole host of species from around the globe. Drawing together evolutionary theory, ecology, population biology, genetics, and physiology and anatomy, it explains the mechanisms and motivations behind a range of animal movements.

The Selfish Gene is a classic exposition of evolutionary thought. In it Professor Dawkins articulates a gene's eye view of evolution - a view giving centre stage to these persistent units of information, and in which organisms can be seen as vehicles for the replication of genes. The book provoked widespread and heated debate, which in part led Dawkins to write The Extended Phenotype, in which he gave a deeper clarification of the central concept of the gene as the unit of selection, as well as contributing his own development of this insight. For the first time, The Extended Selfish Gene brings these two books together, by including two key chapters from The Extended Phenotype. These chapters provide Dawkins's detailed and powerful response to two issues raised by critics of The Selfish Gene: the accusations of genetic determinism (the idea that our behaviour is entirely determined by our genes), and of "adaptationism " (that all traits are indiscriminately perceived to be adaptations resulting from natural selection). While written in particular for the biology community, Dawkins's clarity of expression allows these chapters to be accessible to all who are seriously engaged with the gene's eye view and its implications. The imaginative, powerful, and stylistically brilliant Selfish Gene not only brought the insights of Neo-Darwinism to a wide audience, but galvanized the biology community, generating much debate and stimulating whole new areas of research. Forty years later, its insights remain as significant as on the day it was published. Along with the two extra chapters, The Extended Selfish Gene includes a new epilogue to The Selfish Gene from the author which highlights the relevance of the gene's eye view to evolutionary biology today.

Strategies and Tactics of Behavioral Research and Practice focuses on the most effective methods for measuring and evaluating changes in behavior. The authors provide the rationale for different procedures for measuring behavior and designing within-subject comparisons between control and intervention conditions. The text explains the strengths and weaknesses of methodological alternatives for every topic so that behavioral researchers and practitioners can make the best decisions in each situation. This classic text has been extensively revised to be more accessible and practical. Not only does it feature much more discussion of how research methods are relevant to today's practitioners, it also includes additional examples based on field research and service delivery scenarios. With expanded coverage on creating experimental designs, as well as new chapters on behavioral assessment, the statistical analysis of data, and ethical issues associated with research methods, this book provides a strong foundation for direct behavioral measurement, within-subject research design, and interpretation of behavioral interventions. Enriched with more pedagogical features, including key terms, tables summarizing important points, figures to help readers visualize text, and updated examples and suggested readings, this book is an invaluable resource for students taking courses in research methods. This book is appropriate for researchers and practitioners in behavior analysis, psychology, education, social work, and other social and health science programs that address questions about behavior in research or practice settings.

Animal Learning and Cognition: An Introduction provides an up-to-date review of the principal findings from more than a century of research into animal intelligence. This new edition has been expanded to take account of the many exciting developments that have occurred over the last ten years. The book opens with a historical survey of the methods that have been used to study animal intelligence, and follows by summarizing the contribution made by learning processes to intelligent behavior. Topics include Pavlovian and instrumental conditioning, discrimination learning, and categorization. The remainder of the book focuses on animal cognition and covers such topics as memory, navigation, social learning, language and communication, and knowledge representation. Expanded areas include extinction (to which an entire chapter is now devoted), navigation in insects, episodic memory in birds, imitation in birds and primates, and the debate about whether primates are aware of mental states in themselves and others. Issues raised throughout the book are reviewed in a concluding chapter that examines how intelligence is distributed throughout the animal kingdom. The broad spectrum of topics covered in this book ensures that it will be of interest to students of psychology, biology, zoology, and neuroscience. Since very little background knowledge is required, the book will be of equal value to anyone simply interested in either animal intelligence, or the animal origins of human intelligence. This textbook is accompanied by online instructor resources which are free of charge to departments who adopt this book as their text. They include chapter-by-chapter lecture slides, an interactive chapter-by-chapter multiple-choice question test bank, and multiple-choice questions in paper and pen format.

"Cheetahs of the Serengeti Plains" is the most comprehensive account of carnivore social behavior to date. Synthesizing more than a decade of research in the wild, this book offers a detailed account of the behavior and ecology of cheetahs. Compared with other large cats, and other mammals, cheetahs have an unusual breeding system; whereas lions live in prides and tigers are solitary, some cheetahs live in groups while others live by themselves. Tim Caro explores group and solitary living among cheetahs and discovers that the causes of social behavior vary dramatically, even within a single species.
Why do cheetah cubs stay with their mother for a full year after weaning? Why do adolescents remain in groups? Why do adult males live in permanent associations with each other? Why do adult females live alone? Through observations on the costs and benefits of group living, Caro offers new insight into the complex behavior of this extraordinary species. For example, contrary to common belief about cooperative hunting in large carnivores, he shows that neither adolescents nor adult males benefit from hunting in groups.
With many surprising findings, and through comparisons with other cat species, Caro enriches our understanding of the evolution of social behavior and offers new perspectives on conservation efforts to save this charismatic and endangered carnivore.

As a model organism, the chick has provided valuable insights into broad issues of development in higher animals. The complex interactions between genetic, hormonal and environmental factors which occur in the developing chick provide a potent argument against unitary causal explanations for differences in behaviour. Study of the behaviour of the chick is also relevant to poultry science and the welfare of domesticated birds. This book reviews research on the development of brain and behaviour in the chick and juxtaposes this with similar work on other avian and, to a lesser extent, mammalian species. It begins by outlining the developmental stages of the chick embryo, including the effects of environmental stimulation. Behaviour and the neurochemistry of development and memory formation in the posthatching period are then discussed. The transitions that occur during the first two to three weeks of posthatching life are described, particularly in terms of changing hemispheric dominance. The final chapter examines avian cognition and some issues of welfare for the domestic chicken. The book provides a thorough review of the subject and will interest workers in animal neurophysiology and behaviour, experimental psychologists, and poultry scientists.

Migration is a key process in the population dynamics of many insects, including some of the most damaging pests. Multidisciplinary research into the importance of migration, in recent decades, has produced many new insights. This book reviews current understanding of the ecological, behavioural, physiological and genetic bases of insect migration. The first part describes migration systems in Europe, Asia, Africa, Australia, and America, with an emphasis on the role of regional weather and climate. The second part considers insects adaptation to migration; it covers: aerodynamics and energetics; the integration of migration in insect life cycles; environmental and genetic regulation of migratory potential; and the evolutionary implications of habitat heterogeneity and variability. The book then addresses the application of this knowledge to operational pest forecasting.

This book is the only account of what honeybees actually see. Bees detect some visual features such as edges and colours, but there is no sign that they reconstruct patterns or put together features to form objects. Bees detect motion but have no perception of what it is that moves, and certainly they do not recognize "things" by their shapes. Yet they clearly see well enough to fly and find food with a minute brain. Bee vision is therefore relevant to the construction of simple artificial visual systems, for example for mobile robots. The surprising conclusion is that bee vision is adapted to the recognition of places, not things. In this volume, Adrian Horridge also sets out the curious and contentious history of how bee vision came to be understood, with an account of a century of neglect of old experimental results, errors of interpretation, sharp disagreements, and failures of the scientific method. The design of the experiments and the methods of making inferences from observations are also critically examined, with the conclusion that scientists are often hesitant, imperfect and misleading, ignore the work of others, and fail to consider alternative explanations. The erratic path to understanding makes interesting reading for anyone with an interest in the workings of science but particularly those researching insect vision and invertebrate sensory systems.

The cuckoos are the most variable birds in social behavior and parental care: a few cuckoos are among the most social of all birds and rear their young in a common nest; most cuckoos are caring parents that rear their own young with some females laying a few eggs in the nests of others; while many cuckoo species are brood parasites who leave their eggs in the nests of other birds to rear, with their young maturing to kill their foster nestmates. In The Cuckoos, Robert B. Payne presents a new evolutionary history of the family based on molecular genetics, and uses the family tree to explore the origins and diversity of their behaviour. He traces details of the cuckoos' biology to their original sources, includes descriptions of previously unpublished field observations, and reveals new comparisons of songs showing previously overlooked cuckoo species. Lavishly illustrated with specially commissioned color plates and numerous maps, halftones, and line drawings, The Cuckoos provides the most comprehensive and up-to-date account of this family yet available.

Just over one hundred and thirty years ago Charles Darwin, in The Descent of Man and Selection in Relation to Sex (1871), developed remarkably accurate conclusions about man's ancestry, based on a review of general comparative anatomy and psychology in which he regarded sexual selection as a necessary part of the evolutionary process. But the attention of biologists turned to the more general concept of natural selection, in which sexual selection plays a complex role that has been little understood. This volume significantly broadens the scope of modern evolutionary biology by looking at this important and long neglected concept of great importance.

In this book, which is the first full discussion of sexual selection since 1871, leading biologists bring modern genetic theory and behavior observation to bear on the subject. The distinguished authors consider many aspects of sexual selection in many species, including man, within the context of contemporary evolutionary theory and research. The result is a remarkably original and well-rounded view of the whole concept that will be invaluable especially to students of evolution and human sexual behavior. The lucid authority of the contributors and the importance of the topic will interest all who share in man's perennial fascination with his own history.

The book will be of central importance to a wide variety of professionals, including biologists, anthropologists, and geneticists. It will be an invaluable supplementary text for courses in vertebrate biology, theory of evolution, genetics, and physical anthropology. It is especially important with the emergence of alternative explanations of human development, under the rubric of creationism and doctrines of intelligent design.

Body fluid regulation is pivotal to human health and is served by extensive clinical and pre-clinical science. By combining modern advances with previous findings in the field, this book presents a comprehensive treatment of major experiments, theories, and new advances in the field of body fluid regulation, thirst, and drinking. It features the main integrative brain mechanisms for fluid regulation, the development of such systems, fluid balance during heat and exercise, aging and clinical disorders, and comparative aspects of fluid regulation. The volume focuses on mammalian thirst or drinking behaviour alongside relevant aspects of the physiology of fluid balance. The principal fluid compartments and their regulation by both intakes and losses are highlighted, using both human and animal studies to illustrate the main concepts.

In this first comprehensive synthesis of the literature on food hoarding in animals, Stephen B. Vander Wall discusses how animals store food, how they use food and how this use affects individual fitness, why and how food hoarding evolved, how cached food is lost, mechanisms for protecting and recovering cached food, physiological and behavioral factors that influence hoarding, and the impact that hoarding animals have on plant populations and plant dispersal. He then provides detailed coverage of hoarding behavior across taxa--mammals, birds, and arthropods--to address issues in evolution, ecology, and behavior.
Drawings, photographs, and appendixes document complex and intrinsically interesting food-hoarding behaviors, and the bibliography of nearly 1,500 sources is itself an invaluable and unique reference.

Better known as the "predator-prey relationship," the consumer-resource relationship means the situation where a single species of organisms consumes for survival and reproduction. For example, Escherichia coli consumes glucose, cows consume grass, cheetahs consume baboons; these three very different situations, the first concerns the world of bacteria and the resource is a chemical species, the second concerns mammals and the resource is a plant, and in the final case the consumer and the resource are mammals, have in common the fact of consuming. In a chemostat, microorganisms generally consume (abiotic) minerals, but not always, bacteriophages consume bacteria that constitute a biotic resource. 'The Chemostat' book dealt only with the case of abiotic resources. Mathematically this amounts to replacing in the two equation system of the chemostat the decreasing function by a general increasing then decreasing function. This simple change has greatly enriched the theory. This book shows in this new framework the problem of competition for the same resource.

Mama's Last Hug is a fascinating exploration of the rich emotional lives of animals, beginning with Mama, a chimpanzee matriarch who formed a deep bond with biologist Jan van Hooff. Her story and others like it-from dogs "adopting" the injuries of their companions, to rats helping fellow rats in distress, to elephants revisiting the bones of their loved ones-show that humans are not the only species with the capacity for love, hate, fear, shame, guilt, joy, disgust, and empathy. Frans de Waal opens our hearts and minds to the many ways in which humans and other animals are connected.

Originally published in 1988, this title explores and contrasts means and ends psychology with conventional psychology - that of stimuli and response. The author develops this comparison by exploring the general nature of psychological phenomena and clarifying many persistent doubts about psychology. She contrasts conventional psychology (stimuli and responses) involving reductionistic, organocentric, and mechanistic metatheory with alternative psychology (means and ends) that is autonomous, contextual, and evolutionary.