Thisisnotintendedtobeacomprehensivetextbookofentomology andarachnology, butratheraconcisesynthesisofcertainbasic informationrequiredfor BSc(Hons)andMSc(Entomology) examinations. Theapproachisprimarilyfunctional: forinstance, theskeletalandwaterproofingpropertiesofthearthropodcuticle arediscussed, butnotitsbiochemistry;andIhaveincludedonly thosepointswithwhichIbelievealladvancedstudentsofthesub. jectoughttobefamiliar. Someaspectsaretodayregardedas outdated;othersdonotappearinanycurrenttexts, butIhave includedthembecauseIconsiderthemtobeimportant. Innoway, therefore, shouldthisberegardedasabookofreference. Tobe frank, itconsistsofamassofoversimplificationsandunqualified generalizationswhichareintendedtoclarifythecomplexprinciples underlyingthem. OncetheseprincipleshavebeenthorougWy grasped, thereaderwillhaveacquiredasufficientlybroadapproach tobeabletogetthebestvaluefrommoreadvancedtreatises. MythanksareduetoDrs. JohnDalingwater, AndrewMilner, andespeciallyPaulHillyardfortheiradviceonpalaeontological matters;toProfessorsEinarBursellforpermissiontouseFig. 8, takenfromhis An Introduction to Insect Physiology(Academic Press), NeilF. HadleyandtheEditorsof American Scientistto reproduceFigs. 52,53, andtoF. SchallerforFig. 42fromachap terhewroteinGupta(ed. ) Arthropod Phylogeny(VanNostrand Reinhold). Finally, IwouldliketoexpressmygratitudetoRoy Abrahamsforredrawingthefiguresshowinginsectwingvenations andtoEileenBerghfortypingthemanuscript. Thebookwas completedduringtenureofaLeverhulmeEmeritusFellowship whichIampleasedtoacknowledge, whileProfessorN. A. Mitchison FRSkindlyprovidedmewithaccommodationinUniversityCol. lege, London. J. L. CWUDSLEY-THOMPSON Contents 1 PalaeontologyandPhylogeny. . . . . . . . . . . . . . . . . . 1 1. 1 TheEarliestArthropodanFossils. . . . . . . . . . . . . . . . 1 1. 2 TheFirstTerrestrialArthropods. . . . . . . . . . . . . . . . . 2 1. 3 EvolutionintheArthropods. . . . . . . . . . . . . . . . . . . 4 1. 3. 1 EmbryologicalConsiderations. . . . . . . . . . . . . . . . . . 4 1. 3. 2 ComparativeMorphology. . . . . . . . . . . . . . . . . . . . . 5 1. 3. 3 MonophyleticorPolyphyleticOrigins. . . . . . . . . . . . . 6 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 2 ImplicationsofLiveonLand. . . . . . . . . . . . . . . 10 2. 1 TheSignificanceofSize 10 2. 1. 1 Size, SkeletonsandAllometry 11 2. 1. 2 AllometricGrowth 12 2. 2 WaterRelations 12 2. 3 TheConquestoftheLand 14 2. 4 . TheIntegument 14 2. 4. 1 TheEndocuticle 15 2. 4. 2 TheExocuticle 15 2. 4. 3 TheEpicuticle 16 2. 5 GrowthandEcdysis 18 2. 6 Respiration: Lung-BooksandTracheae 19 2. 7 NutritionandExcretion 21 2. 7. 1 Nutrition 21 2. 7. 2 Excretion 22 2. 8 EcologicalConsiderationsofSize 23 FurtherReading 25 3 TheConquestoftheLandbyCrustacea. . . . . . . . . . . . 26 3. 1 TypesofAdaptation. . . . 26 3. 2 TransitionfromWatertoLandinAmphipoda 26 3. 3 TransitionfromWatertoLandinDecapoda 28 3. 4 TransitionfromWatertoLandinIsopoda 30 3. 4. 1 Morphology 30 3. 4. 2Physiology. . . . . . . . . . . . . . . . . . . . . . . . . . . 31 3. 4. 3 Behaviour 34 VIII Contents 3. 5 Conclusion 35 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . 36 4 InsectPhylogenyandtheOriginofFlight. . . . 37 4. 1 AncestryofInsects 37 4. 2 TheOriginofWings 38 4. 2. 1 Apte10taandtheAncestryofSpiders 39 4. 3 Paranota1Theory 40 4. 4 TrachealGillTheory . 42 4. 5 SelectionforFlight 43 4. 6 PhylogenyoftheLowerInsectOrders . 44 4. 6. 1 Fossil

The nemerteans are a fascinating, common and often locally abundant group of invertebrates, yet for long have attracted the attention of only a handful of scientists. In recent years, however, increasing numbers of people have developed diverse research interests in the group with the result that our knowledge of these worms has rapidly advanced. Clearly there is a need for a regular series of international meetings where individuals working on these animals can meet to exchange ideas, review developments concerning nemertean biology and freely discuss future proposals. The first such meeting was held in Philadelphia during December 1983. Twenty-seven scientists from eight countries participated in the Second International Meeting on Nemertean Biology, held at the Tjarno Marine Biological Laboratory, Sweden, 11-15 August 1986. The meeting was divid ed into five sessions: two dealt with ecological studies, two with nemertean taxonomy, and the final session covered aspects of general biology. A total of 26 papers was presented; four, by N. Anadem, G. Berg, 1. Bierne, and 1.M. TurbeviIIe, were for different reasons not submitted for inclusion in this volume. Three additional presentations were made on behalf of B. Kulikova, E. N. WiIImer, and Z. Yin, all of whom were unable to participate in the meeting.

Bees existed long before human beings, but our future is perhaps more reliant upon them than any other species. They pollinate 80 per cent of the world's crops and plants, but how much do we really know about them? Small, clever and mysterious, the honeybee in particular has long been celebrated in human culture as a sacred insect, a symbol of the sun, bridging the gap between our world and the next. They are expert communicators, skilled aviators and natural alchemists, turning fresh nectar into sweet, golden honey. They are also in trouble and need our help. This beautifully illustrated guide explores the honeybee's historic relationship with humans, the basics of beekeeping, and how we can help save the bees' dwindling population.

A rapidly growing interdisciplinary field, disease ecology merges key ideas from ecology, medicine, genetics, immunology, and epidemiology to study how hosts and pathogens interact in populations, communities, and entire ecosystems. Bringing together contributions from leading international experts on the ecology of diseases among invertebrate species, this book provides a comprehensive assessment of the current state of the field. Beginning with an introductory overview of general principles and methodologies, the book continues with in-depth discussions of a range of critical issues concerning invertebrate disease epidemiology, molecular biology, vectors, and pathogens. Topics covered in detail include: Methods for studying the ecology of invertebrate diseases and pathogens Invertebrate pathogen ecology and the ecology of pathogen groups Applied ecology of invertebrate pathogens Leveraging the ecology of invertebrate pathogens in microbial control Prevention and management of infectious diseases of aquatic invertebrates Ecology of Invertebrate Diseases is a necessary and long overdue addition to the world literature on this vitally important subject. This volume belongs on the reference shelves of all those involved in the environmental sciences, genetics, microbiology, marine biology, immunology, epidemiology, fisheries and wildlife science, and related disciplines.

This major reference work contains essential information on arthropod-borne infections affecting humans and domesticated animals. The encyclopedia is a key reference source for anyone working in medical and veterinary science, and related fields. Features of "iThe Encyclopedia of Arthropod-transmitted Infections" are 150 entries, describing arboviral, viral, bacterial and rickettsial, spirochaetal, protozoal and filarial infections, and the vectors that transmit themInformation on disease distribution, clinical symptoms, diagnosis, transmission cycles, vector life-cycles, and treatment and control measures. Figures, tables and photographs illustrate the text. Following each entry is a selected bibliography, to aid further reading on the topic. Over 80 different international authors, with expertise in medicine, veterinary science, parasitology, entomology, epidemiology, microbiology, and zoology have contributed to the encyclopedia

Proceedings of the 6th International Symposium on Insect-Plant Relationships (Pau 1986)

Palms constitute one of the largest botanical families, and include some of the world s most important economic plants. They are also unequalled as outdoor and indoor ornamental plants, and include many species that are essential components of the ecosystems of tropical and other warm regions. This book reviews the inter-relationships between palms and insects, emphasizing the similarities in different world regions. The host plants, distribution, and bionomics of representative insects are discussed according to their feeding sites on palms (foliage, flowers, fruits, and stems) and their taxonomic groups. Host and distribution records for the most extensively represented insect families on palms are tabulated. Pest management and field techniques are also covered. This book is recommended reading for tropical biologists and agriculturalists, including entomologists, horticulturists and tropical ecologists as well as palm nursery growers, managers and enthusiasts."

Few branches of the biological sciences have developed to such an extent as has ecology in the recent decades. The successful development of this science is directly related to need to create a sound scientific basis by which we can control our diminishing natural resources and integrate this with the control of biological systems of the component species. Studies on the bioenergetics of ecosystems, as well as on the home ostatic mechanisms functioning at the populations and biocenoses levels are of great importance in this respect. The results of these studies are very significant in forestry management which deals with multiannual tree communities - forest stands. It is particularly important in this of the necessity for the possible respect for economic planning, in view prolonged maintenance of forest biocenoses as stable systems. Neither in the present nor in the immediate future can be protection of forest ecosystems consisting of natural plant and animal communities be achieved by the cultivation of resistant forms or the intensification of chemical intervention."

The behaviour of these bees has been adapted to some extent to meet beekeeping requirements but few advances have been made in altering their behaviour so that honey and wax production is increased. Bee pheromone research aims to identify the pheromones and make analogous synthetic chemicals thus allowing the control of the colony's activities and enhancing honey production. This book evaluates the present state of knowledge of each known pheromone, discusses the pheromones of the stingless bees and describes the effect of pheromones on the social organization and mating behaviour of bumblebees. Suggestions are made for further and different types of research into pheromones. The book should be of interest to zoologists, entomologists and general biologists.

The nature and diversity of presentations at the second International Conference on Insect Neurochemistry and Neurophysiology (ICINN--86) held at the University of Maryland on August 4-6. 1986, attest to the vital ity and broad scope of research in insect neuroscience. The present vol ume is a written account of the invited lectures, contributed papers, and posters presented at the conference, and as such, serves as a fair indica tor of the trends in current research in this field here and abroad. The principal portion of this book consists of seven review papers that were presented by invited speakers. Although the topics vary wide ly, they reflect on and emphasize the main theme of the conference, i. e., the nature and function of molecular messengers that communicate be meen the central nervous system and organs or tissues involved in the growth, development, reproduction, and behavior of insects. This empha sis is continued in the following three sections on neurochemistry, neuro physiology, and neuroanatomy, although no conscious effort was made by the organizers to highlight these particular fields of neuroscience. It is evident that the recent advances in both physical and chemical analyti cal techniques have made possible the acquisition of structurally defined probes, the long sought-after tools for unraveling the secrets of endogen ous communication. Each section of short papers derived from the oral and poster presentations at the conference is prefaced by an overview that highlights and summarizes the section's content."

Beetles, the most diverse group of insects, are often abundant in terrestrial and freshwater ecosystems. Many species are under threat from human changes to natural environments, and some are valuable tools in conservation, because they respond rapidly to changes that occur. Knowledge of these responses, of both abundance and composition of assemblages, enable use of some beetles to monitor environmental changes. Beetles impinge on humanity on many ways: as cultural objects, desirable collectables, major pests and competitors for resources need by people, as beneficial consumers of other pests, and by ensuring the continuity of vital ecological processes. This book is the first major global overview of the importance of conservation of beetles, and brings together much hitherto scattered information to demonstrate the needs for conservation, and how it may be approached. It is a source of value to students, research workers, conservation biologists and ecosystem managers as an introduction to the richness and importance of this predominant component of invertebrate life.

Proceedings of the 2nd Symposium held at Zvikovske podhradi, September 2-8, 1984

The increasing use of integrated crop management, often requiring a reduction in the reliance upon chemical control, means that the need to rapidly identify pest nematodes has never been greater. This second edition of this standard reference work familiar to all plant nematologists is therefore even more useful than its predecessor published in 1986. The in-depth description of the life histories of the genera of the Tylenchida have been retained and brought up-to-date through the inclusion of all the research carried out between the publication of the last edition and this new edition. This expanded edition includes detailed diagnoses of well over 200 genera and familial and ordinal groups and the book is heavily illustrated with drawings of type or representative species. These, together with comprehensive lists of species and genera and their synonymies provide the foundation for the status and validation of each taxon within the Tylenchida. A considerable amount of information is provided regarding the biology, ecology and pathogenicity of these parasites. This book is the only comprehensive reference work on this important group of nematodes.

Bark Beetle Management, Ecology, and Climate Change provides the most updated and comprehensive knowledge on the complex effects of global warming upon the economically and ecologically important bark beetle species and their host trees. This authoritative reference synthesizes information on how forest disturbances and environmental changes due to current and future climate changes alter the ecology and management of bark beetles in forested landscapes. Written by international experts on bark beetle ecology, this book covers topics ranging from changes in bark beetle distributions and addition of novel hosts due to climate change, interactions of insects with altered host physiology and disturbance regimes, ecosystem-level impacts of bark beetle outbreaks due to climate change, multi-trophic changes mediated via climate change, and management of bark beetles in altered forests and climate conditions. Bark Beetle Management, Ecology, and Climate Change is an important resource for entomologists, as well as forest health specialists, policy makers, and conservationists who are interested in multi-faceted impacts of climate change on forest insects at the organismal, population, and community-levels.

Insects are the most abundant and diverse organisms that inhabit our planet and are found in all the world's forest ecosystems. Many feed and/or breed on parts of trees. Some perform important functions, such as pollination or break-down of dead vegetation. Others weaken, deform or kill trees, and compete with humans for the many goods and services that trees and forests provide. "Forest" "Entomology: A Global Perspective" examines forest insects in a global context and reviews their dynamics, interactions with humans and methods for monitoring and management of species that damage forests. Also provided are 235 profiles of forest insects, worldwide. A series of tables provides summaries of the distribution and hosts of many more species. Included are those that damage forests, others that are simply curiosities and some that are beneficial. This book is designed as a reference for students, practicing foresters and forest health specialists, especially for those who work internationally or are concerned with species that have the potential to expand their ranges via international trade, travel or environmental changes.

This text provides an introductory account of the physiology of insects, intelligible to any reader with an elementary knowledge of biology. It should be of interest to undergraduate students of biology, entomology and agriculture.