About 95 per cent of all known animal species are invertebrates. A knowledge of their sexual, reproductive, and development biology is essential for the effective management of species that are economically useful to man or are harmful to him, his crops, and livestock. This treatise is the first to cover all aspects of reproduction and development of the entire spectrum of invertebrates -- terrestrial, marine, freshwater, brackish-water, free-living, and parasitic. The chapters, by leading world experts in their fields, are up-to-date and informative, and suggest a number of problems for future research. "Progress in Male Gamete Ultrastructure and Phylogeny" (issued in parts A--C) is the ninth volume in the series.
Contents Preface to the Progress Series; Preface to Volume IX, Part A; Contributors; Porifera, N. Boury-Esnault and B.G.M. Jamieson; Cnidaria and Ctenophora, Peter L. Harrison and B.G.M. Jamieson; Platyhelminthes, Nikki A. Watson; Nemertea, Ake Franzen and Bjorn A. Afzelius; Rotifera, Giulio Melone and Marco Ferraguti; Gastrotricha, Maria Balsamo, Elena Fregni and Marco Ferraguti, Kinorhyncha, Andrey V. Adrianov and Vladimir V. Malakhov; Nematomorpha, Roberto Valvassori, Magda de Eguileor, Annalisa Grimaldi and Giulio Lanzavecchia; Acanthocephala, Marcella Carcupino and Bahram S. Dezfuli; Subject Index; Species Index.

Biomonitoring is a vital and rapidly growing field. Freshwater Biomonitoring and Benthic Macroinvertebrates presents a state-of-the-art look at the use of benthic macroinvertebrates (aquatic insects, molluscs, crustaceans, and worms) in the biological assessment of water quality in lakes and streams. The use of these organisms has increased dramatically in the past two decades in both North America and Europe; no other comprehensive overview of the topic is available. This book contains contributions from nineteen experts from North America, Europe, and Australia. Some chapters contain updates and syntheses of new information on previously reviewed topics, while others present the first detailed coverage of a topic. The book includes much useful reference material on the history of biomonitoring with invertebrates as well as work of a more practical nature. The design, implementation, analysis and interpretation of benthic surveys are discussed in detail, as are toxicity testing and field experiments. This timely and important book will be of interest to professional entomologists, invertebrate zoologists, and aquatic ecologists, especially those in applied areas concerned with environmental quality, preservation, and restoration. Those working in the growing environmental biotechnology field, or in government agencies responsible for monitoring drinking water, environmental quality, or marine habitats, will also find many new ideas here.

Insect Learning is a comprehensive review of a new field. Until recently, insects were viewed as rigidly programmed automatons; now, however, it is recognized that they can learn and that their behavior is plastic. This fundamental change in viewpoint is causing a re-examination of all aspects of the relationship between insects and their environment. This change in perspective is occurring at a time of heightened interest in brain function in both vertebrates and invertebrates. Insects potentially play a major role in this expanding area. Because of their experimental tractability and genetic diversity, they provide unique opportunities for testing hypotheses on the ecology and evolution of learning. As organisms of economic importance, they are perennial objects of research by both basic and applied scientists. Insect Learning covers both social and non-social insects from multiple perspectives. The book covers mechanisms; syntheses of work on physiology, behavior, and ecology; and micro- and macroevolution. The concluding section discusses future directions for research, including applications to pest management.

Sex ratio, the proportion of a progeny that is female, is an extremely important fitness trait. It is particularly interesting in relation to haplodiploidy, a variety of parthenogenesis in which organisms reproduce without the benefit of fertilization. This study arrives at conclusions drawn from new empirical studies: that biased sex ratios are characteristic of haplodiploid species; that these species are characteristically colonizing species with genetics suited both to spatially and temporally unpredictable environments; that manipulation of haplodiploid biological control agents, or pests that are to be controlled, depends on understanding the determinants of sex ratio; and that because evolutionary theory predicts that haplodiploids have the capacity to evolve faster than diploid organisms, haplodiploid species are the organisms of choice in biological control strategies.

Insect Chemical Ecology provides a comprehensive view of how natural selection acts upon interacting organisms and how particular physical and biological properties of chemical compounds act as constraints upon which natural selection may act. Individual chapters raise specific questions as to the nature of these interactions. The first part contains reviews on antagonistic and mutualistic chemical interactions, the raw materials' of chemical evolution, the economics of offensive and defensive chemicals, and neurobiology. The second part discusses particular problems such as the evolution of resistance, insect pollination, learning, pheromones, sequestration of semiochemicals, the role of microorganisms, sex attractants, the evolution of host races and biotypes, and the role of semiochemicals and the evolution of sociality of insects. The last chapter discusses the role of chemical-based pest management programs in an ecological and evolutionary framework.

Insect Chemical Ecology provides a comprehensive view of how natural selection acts upon interacting organisms and how particular physical and biological properties of chemical compounds act as constraints upon which natural selection may act. Individual chapters raise specific questions as to the nature of these interactions. The first part contains reviews on antagonistic and mutualistic chemical interactions, the raw materials' of chemical evolution, the economics of offensive and defensive chemicals, and neurobiology. The second part discusses particular problems such as the evolution of resistance, insect pollination, learning, pheromones, sequestration of semiochemicals, the role of microorganisms, sex attractants, the evolution of host races and biotypes, and the role of semiochemicals and the evolution of sociality of insects. The last chapter discusses the role of chemical-based pest management programs in an ecological and evolutionary framework.

Harold Maxwell-Lefroy, the founder of Rentokil, was a maverick and a man of enormous drive and energy. From an early age he was fascinated with the insect world, and his thorough understanding of species' life cycles and habits, in its practical application, was to change the face of agriculture in several parts of the world. He was among the first really to apply the scientific method to dealing with insect pests, and the agriculture of the Caribbean and India still owes him an enormous debt. His book Indian Insect Pests is still in print, an invaluable resource to Indian agriculturalists. In the Caribbean he saved the sugar crop which had been ravaged by pests, and was then sent to India as the official entomologist. Here his energy and drive led to an education programme for Indian farmers that for the first time showed them that the devastating consequences of insect pests were avoidable, along with the destruction of livelihoods that had always been an occupational hazard. He became the first Professor of Entomology at Imperial College and developed patented anti-pest chemical treatments that led him to create Rentokil towards the end of his life - trademark rules barred him from calling it Entokil, as he had wanted to. He went on to save the roof of Westminster Hall from the death-watch-beetle infestation that would certainly have led to its collapse. But he was also an inveterate risk-taker, who drove without regard for his own safety, and applied the same principles to his scientific practice. He died at the young age of 48, overcome by the poisonous gases he was developing - without the proper breathing equipment. Rentokil is his most tangible legacy, but it all began with one man's single-minded dedication to the application of science.

The acridoid grasshoppers in general and the various species of swarm-forming locusts in particular have been among the most favoured subjects of insect physi ology, behaviour and ecology for many years. Several factors contribute to this popularity: their abundance in nature and their ease of culture in the laboratory, their relatively large size, and most of all, perhaps, their agricultural importance and the consequent intermittent availability of funds for their study. These factors together have inspired a large amount of experimental work, and this in tum has produced a new and often over-riding reason for working on acridoids -the huge body of available background information and know-how that has built up about these insects. This state of affairs is well seen in insect neurobiology. Only a restricted number of insect types are commonly used in this discipline, and originally most of them were selected for reasons of convenience and availability: grasshoppers, cock roaches, crickets, flies, bees and moths are the most important. Each of these in sects is the subject of the attentions of one or more major groups of neurobiolo gists, but neurobiological articles on acridoids probably exceed in number those on all other insects combined, at least if articles on the molecular biology of the nervous system of Drosophila are excluded.

The search for new strategies of pest control with safer molecules is currently of great importance and interest. Microbe-mediated biological crop protection is an attractive and promising technology with no concern for a negative impact on the environment and biodiversity. Microbial hydrolytic enzymes such as proteases, chitinases, lipases, etc. are attractive for this purpose. They present toxic properties and act synergistically to control pest attacks. Also, some metabolites, that microorganisms produce for their survival or defense, can be explored and exploited for plant protection. The focus of this Volume is on the potential of microbial hydrolytic enzymes and their metabolites in agroecosystem functioning. Subsequent chapters review topics such as microbial hydrolytic enzymes as powerful management tools, chitinases in IPM of agro-horticultural crops, metabolites as pesticides and the importance of the metabolites of entomopathogenic fungi, metabolites and virulence factors. Other topicas include: microbial-based nanoparticles, recombinant DNA technologies to improve the efficacy of microbial insecticides, the effects of entomopathogens on insect predators and parasitoids, and the management of major vegetable insect pests. This Volume provides detailed accounts on the safe use of microbial products for sustainable management of insect pests. Its aim is to build solid foundations for the students, teachers, and researchers interested in eco-friendly management of important insect crop pests.

The Biological Sciences are in the midst of a scientific rev olution. During the past decade under the rubric of molecu lar biology, chemistry and physics have assumed an integral role in biological research. This is especially true in ge netics, where the cloning of genes and the manipulation of genomic DNA have become in many organisms routine laboratory procedures. These noteworthy advances, it must be empha sized, especially in molecular genetics, are not autonomous. Rather, they have been accomplished with those organisms whose formal genetics has been documented in great detail. For the beginning student or the established investigator who is interested in pursuing eukaryote molecular genetic re search, Drosophila melanogaster, with its rich body of formal genetic information is one organism of choice. The book "Drosophila Genetics. A Practical Course" is an indispens able source of information for the beginner in the biology and formal genetics of Drosophila melanogaster. The scope of this guide, a revision and enlargement of the original German language version, is broad and instructive. The information included ranges from the simple, but necessary, details on how to culture and manipulate Drosophila flies to a series of more sophisticated genetic experiments. After completing the experiments detailed in the text, all students - neophyte or experienced - will be richly rewarded by having acquired a broad base of classical genetics information relevant for the biologist in its own right and prerequisite to Drosophila genetics research - formal and/or molecular. Davis, California, Melvin M."

"Vincent Dethier shows us how to listen for sound in fields, edges, and woods and to become aware of the movements that accompany sound...We learn from his sounds what kind of person, capable of this kind of interest and care, is attending to our minds. His own sound becomes part of the community of sound common to most, if not nearly all, life, so we are doubly trained to hear, and we become doubly committed to understanding and caring for all forms of life." --A. R. Ammons, from the foreword

The use of hermit crabs as an experimental model for motivational studies is discussed in this book by expert behavioural ecologists. The book is timely since there is a growing recognition that the integration of causal and functional studies are needed, which is discussed by the authors. This book should be of interest to a wide range of behavioural ecologists.

In this book, contributors focus on the reproductive behaviour of insects. The topics they discuss range from mate finding, host selection and oviposition behaviour to the competition for resources and the behaviour of feeding larvae. The papers reflect both a diversity of approach and differing degrees of conviction as to the relevance of evolutionary theory to the study of insect population. The aim of the book is to encourage the reader to think more critically about the evolution of behaviour, and in particular about the importance of small differences in individual behaviour within natural populations.

In August 1988. the Sixth International Coral Reef Symposium was held in Townsville resulting in an influx of most of the world's coral reef sCientists to the city. We seized this opportunity at the Australian Institute of Marine Science to run a small workshop immediately before the symposium on the outbreaks of the crown-of-thorns starfish. Aeanthaster planei. We invited that small band of mathematicians who had been modelling the phenomenon, (and who may not have normally attended an international meeting so thoroughly dedicated to natural science) to meet with those SCientists who had been been actively working on the phenomenon in the field. John Casti notes in his delightful new book Alternate Realities (Wiley, 1989): 'If the natural role of the experimenter is to generate new observables by which we know the processes of Nature, and the natural role of the mathematician is to generate new formal structures by which we can represent these processes. then the system SCientist finds his niche by serving as a broker between the two. ' I think our book shows the fruits of that brokerage through the wide range of models explored within its pages. the high level of collaboration and interaction across disciplines evident in the individual papers, and in the emerging synthesis that reflects a far deeper understanding of this complex phenomenon than was possible even a few years ago.

This book considers research on insect acoustic behaviour from two divergent approaches to insect behaviour. The first approach involves the current rediscovery of sexual selection and involves the testing of many hypotheses concerning the relative importance of different behavioural strategies to the fitness of the individual. The other approach considers the role of specific neurones within the insect's auditory system. This book aims to integrate these aspects of evolutionary biology and neurophysiology. The book discusses the role of sound in the life and life-cycle of insects and includes a discussion of mate recognition, chorusing, calling and listening.

The first book on entomophagy written in this manner, Edible Insects of the World is an enumeration of 2,141 species of edible insects. For the reader's convenience, insect names are arranged much like a dictionary, first by taxonomic group and then by country. In addition to taxonomic and location information, entries contain applicable details about the history of entomophagy, collecting, cooking, and medicinal uses.

In 1980, the International Symposium on the Ecology of Bruchids Attacking Legumes (Pulses), organized by Dr. Labeyrie, was held at Tours, France. Since then, there has been tremendous progress in the area of Bruchid and Legume research. At the same time, as we face the problems of world-wide population explosion and food shortage, the importance of legumes as the world's major protein source is rapidly increasing, especial ly in tropical regions. Thus, it seemed appropriate to hold the Second Symposium in order to review the recent progress in the control of Bruchids and in the biology and ecology of Bruchids and legumes. This is an important part of the search for ways to integrate these fields with a common perspective. The Second International Symposium on Bruchids and Legumes (ISBL-II) was held in September 6-9, 1989 at Okayama, Japan under the joint auspices of the Japanese Society of Applied Entomology and Zoology and of the Foundation for Advancement of International Science. Significant contributions have originated in Japan on the study of Bruchid and legumes. Most notably, the study on population ecology by Professor S."

The papers included in this volume were amongst day running of the conference, and Ms Julia those presented at the 5th International Epheme Reed, Ms Kim James, Ms Anne Devereaux, roptera Conference and the 9th International Mr Peter Green, Ms Catriona Smith who assist Plecoptera Conference at the Marysville Hotel, ed them, Mr David Ginn and all the staff at the Marysville Australia from the 18th to the 24th of Marysville Hotel also deserve special thanks for February 1987. The conference was attended by the the superb catering and relaxed atmosphere 62 participants from 21 countries. This was the they created which contributed so much to the frrst time the two conferences had been held conference. together, and the frrst time either had been held in All the papers included in this volume were the southern hemisphere. refereed prior to acceptance, and I would like to The papers included here cover a broad spectrum thank the following referees: Dr J. Davis, of research into the two orders of aquatic insects, Dr L. Barmuta, Dr R. Marchant, Mr T. Doeg, with the emphasis on life histories, which was the Dr P. Bailey, Dr S. Bunn, Dr R. Rowe, Dr R. theme of a joint symposium held during the con Pearson, Ms C. Yule and Dr P. Suter. Ms Sue ference. The paper by Dr Brittain was presented Mitchell assisted with the typing and Ms Kerrle as a keynote address to that symposium. The Swadling with the proof-reading of the text."

Spider Evolution: Genetics, Behavior, and Ecological Influences provides a thorough exploration of the evolutionary trail of arachnids, particularly spider species, from prehistoric origins to current sustainability issues. This book analyzes extinct organisms in the Arachnida class, specifically looking at their phylogenomics and molecular footprints to understand evolutionary changes in diversification in today's species. Sections cover spider origins and their influences on behavioral traits, physiology of sensory organs, and biomechanics, also touching on spiders as prey and predators and how their roles have changed in the 400 million years of Arachnida existence. The book then focuses upon current environmental issues facing spider species and how these have, and can, affect the evolution of these organisms. Topics include biodiversity minimization, climate change and natural disasters. This book is a much-needed resource for entomologists and arachnid- or arthropod-driven researchers. Advanced undergraduate and graduate students will also benefit from the historic review, current assessment and future predictions of spider evolution provided in this book.

Covering 100 years of zoological research, the Handbook of Zoology represents a vast store of knowledge. Handbook of Zoology provides an in-depth treatment of the entire animal kingdom covering both invertebrates and vertebrates. It publishes comprehensive overviews on animal systematics and morphology and covers extensively further aspects like physiology, behavior, ecology and applied zoological research. Although our knowledge regarding many taxonomic groups has grown enormously over the last decades, it is still the objective of the Handbook of Zoology to be comprehensive in the sense that text and references together provide a solid basis for further research. Editors and authors seek a balance between describing species richness and diversity, explaining the importance of certain groups in a phylogenetic context and presenting a review of available knowledge and up-to-date references. New contributions to the series present the combined effort of an international team of editors and authors, entirely published in English and tailored to the needs of the international scientific community. Upcoming volumes and projects in progress include volumes on Annelida (Volumes 1-3), Bryozoa, Mammalia, Miscellaneous Invertebrates, Nannomecoptera, Neomecoptera and Strepsiptera and are followed later by fishes, reptiles and further volumes on mammals. Background The renowned German reference work Handbook of Zoology was founded in the 1920's by Professor Willi Kukenthal in Berlin and treated the complete animal kingdom from single cell organisms to mammals in eight thematic volumes: Volume I Protozoa, Porifera, Colenteratea, Mesozoa (1925); Volume II Vermes (1933/34); Volume III Arthropoda ex. Insecta (1927/1932); Volume IV Arthropoda: Insecta; Volume V Solenogastres, Mollusca, Echinoderma (1925); Volume VI Pisces / Amphibia (1930); Volume VII Reptilia / Aves (1931); Volume VIII Mammalia. The Volumes IV Arthropoda: Insecta and VII Mammalia continued publication into the present with the most recent contributions in English language. Adapting to the accelerating speed of scientific discovery in the past decades the Handbook of Zoology entered a next phase in 2010. In the new edition of the Handbook of Zoology, the original eight thematic volumes gave way for smaller and more flexible groupings that reflect the current state of phylogenetic knowledge. All subsequent volumes were published in print as well as e-book format. The Handbook of Zoology is additionally offered as a database, the Handbook of Zoology Online, which can easily be searched and rapidly updated. Original Handbook material (ca. 28 000 pages) has been reordered along taxonomic (instead of bibliographical) categories and forms the historical basis of this Online Reference Work. As a living Online Reference, the content is continuously updated and new content added. The material can be accessed through taxonomic and subject categories as well as free text, with a diversity of linking and search options. Faster publication times through online-first publication, reference- and cross-linking, and make the Handbook of Zoology highly attractive to both authors and users.

The present edition may be regarded as a descendant, much changed and greatly enlarged, of the late Dr A. D. Imms' Outlines of Entomology, first published in 1942. This went through three further editions without much change, but after the death of the original author a fifth, revised edition by Professor O. W. Richards and myself appeared in 1959 and a sixth in 1978. The book now appears in a considerably extended version in which I have tried to provide a more balanced introduction to the whole field of modern entomology by dealing with several aspects of the subject not discussed at all in previous editions. Thus, in addition to innumerable lesser changes in the sections on insect structure, function, development, classification and phylogeny, I have completely recast the earlier chapter on some important modes of life in insects. This now includes a far wider range of biological topics well exemplified by the insects and should, I hope, appeal not only to those already dedicated to entomology but also to others with more general biological interests. A completely new chapter on the biology of insect populations has also been added and may serve to indicate the debt which modern ecological theory owes to work on insect populations. It should hardly be necessary to apologize for introducing a certain amount of elementary mathematics into this account of a subject which is now among the most highly quantitative of biological disciplines.

This volume provides an abundance of valuable first-hand information about the diversity, biology, ecology culture of the portunid crabs of the word seas. Marine crabs play an important role directly or indirectly in the livelihood of millions of people around the world. They have been reported to make up about 20% of all marine crustaceans caught, farmed, and consumed worldwide. Among these marine crabs, portunid crabs (or swimming crabs of the family Portunidae) assume greater significance in the marine industry, owing to their delicate meat with nutritional qualities. Although several species of portunid crabs are edible and commercially important, only a few species have been widely cultivated. This is largely due to the lack of information on the biology of portunid crabs. Keeping this in view, this volume presents the biology and aquaculture of marine portunid crabs. This volume will be of great use for researchers and students of disciplines such as fisheries science, marine biology, aquatic biology and fisheries, and zoology, and will also serve as a standard reference for college, university, and research libraries around the world.

Thisisnotintendedtobeacomprehensivetextbookofentomology andarachnology, butratheraconcisesynthesisofcertainbasic informationrequiredfor BSc(Hons)andMSc(Entomology) examinations. Theapproachisprimarilyfunctional: forinstance, theskeletalandwaterproofingpropertiesofthearthropodcuticle arediscussed, butnotitsbiochemistry;andIhaveincludedonly thosepointswithwhichIbelievealladvancedstudentsofthesub. jectoughttobefamiliar. Someaspectsaretodayregardedas outdated;othersdonotappearinanycurrenttexts, butIhave includedthembecauseIconsiderthemtobeimportant. Innoway, therefore, shouldthisberegardedasabookofreference. Tobe frank, itconsistsofamassofoversimplificationsandunqualified generalizationswhichareintendedtoclarifythecomplexprinciples underlyingthem. OncetheseprincipleshavebeenthorougWy grasped, thereaderwillhaveacquiredasufficientlybroadapproach tobeabletogetthebestvaluefrommoreadvancedtreatises. MythanksareduetoDrs. JohnDalingwater, AndrewMilner, andespeciallyPaulHillyardfortheiradviceonpalaeontological matters;toProfessorsEinarBursellforpermissiontouseFig. 8, takenfromhis An Introduction to Insect Physiology(Academic Press), NeilF. HadleyandtheEditorsof American Scientistto reproduceFigs. 52,53, andtoF. SchallerforFig. 42fromachap terhewroteinGupta(ed. ) Arthropod Phylogeny(VanNostrand Reinhold). Finally, IwouldliketoexpressmygratitudetoRoy Abrahamsforredrawingthefiguresshowinginsectwingvenations andtoEileenBerghfortypingthemanuscript. Thebookwas completedduringtenureofaLeverhulmeEmeritusFellowship whichIampleasedtoacknowledge, whileProfessorN. A. Mitchison FRSkindlyprovidedmewithaccommodationinUniversityCol. lege, London. J. L. CWUDSLEY-THOMPSON Contents 1 PalaeontologyandPhylogeny. . . . . . . . . . . . . . . . . . 1 1. 1 TheEarliestArthropodanFossils. . . . . . . . . . . . . . . . 1 1. 2 TheFirstTerrestrialArthropods. . . . . . . . . . . . . . . . . 2 1. 3 EvolutionintheArthropods. . . . . . . . . . . . . . . . . . . 4 1. 3. 1 EmbryologicalConsiderations. . . . . . . . . . . . . . . . . . 4 1. 3. 2 ComparativeMorphology. . . . . . . . . . . . . . . . . . . . . 5 1. 3. 3 MonophyleticorPolyphyleticOrigins. . . . . . . . . . . . . 6 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 2 ImplicationsofLiveonLand. . . . . . . . . . . . . . . 10 2. 1 TheSignificanceofSize 10 2. 1. 1 Size, SkeletonsandAllometry 11 2. 1. 2 AllometricGrowth 12 2. 2 WaterRelations 12 2. 3 TheConquestoftheLand 14 2. 4 . TheIntegument 14 2. 4. 1 TheEndocuticle 15 2. 4. 2 TheExocuticle 15 2. 4. 3 TheEpicuticle 16 2. 5 GrowthandEcdysis 18 2. 6 Respiration: Lung-BooksandTracheae 19 2. 7 NutritionandExcretion 21 2. 7. 1 Nutrition 21 2. 7. 2 Excretion 22 2. 8 EcologicalConsiderationsofSize 23 FurtherReading 25 3 TheConquestoftheLandbyCrustacea. . . . . . . . . . . . 26 3. 1 TypesofAdaptation. . . . 26 3. 2 TransitionfromWatertoLandinAmphipoda 26 3. 3 TransitionfromWatertoLandinDecapoda 28 3. 4 TransitionfromWatertoLandinIsopoda 30 3. 4. 1 Morphology 30 3. 4. 2Physiology. . . . . . . . . . . . . . . . . . . . . . . . . . . 31 3. 4. 3 Behaviour 34 VIII Contents 3. 5 Conclusion 35 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . 36 4 InsectPhylogenyandtheOriginofFlight. . . . 37 4. 1 AncestryofInsects 37 4. 2 TheOriginofWings 38 4. 2. 1 Apte10taandtheAncestryofSpiders 39 4. 3 Paranota1Theory 40 4. 4 TrachealGillTheory . 42 4. 5 SelectionforFlight 43 4. 6 PhylogenyoftheLowerInsectOrders . 44 4. 6. 1 Fossil

The nemerteans are a fascinating, common and often locally abundant group of invertebrates, yet for long have attracted the attention of only a handful of scientists. In recent years, however, increasing numbers of people have developed diverse research interests in the group with the result that our knowledge of these worms has rapidly advanced. Clearly there is a need for a regular series of international meetings where individuals working on these animals can meet to exchange ideas, review developments concerning nemertean biology and freely discuss future proposals. The first such meeting was held in Philadelphia during December 1983. Twenty-seven scientists from eight countries participated in the Second International Meeting on Nemertean Biology, held at the Tjarno Marine Biological Laboratory, Sweden, 11-15 August 1986. The meeting was divid ed into five sessions: two dealt with ecological studies, two with nemertean taxonomy, and the final session covered aspects of general biology. A total of 26 papers was presented; four, by N. Anadem, G. Berg, 1. Bierne, and 1.M. TurbeviIIe, were for different reasons not submitted for inclusion in this volume. Three additional presentations were made on behalf of B. Kulikova, E. N. WiIImer, and Z. Yin, all of whom were unable to participate in the meeting.