This is the third edition of this popular text which reviews all aspects of earthworm biology and ecology. The book has been updated throughout and extensive additions made. These include a greatly expanded treatment of earthworm community ecology, interactions between earthworms and microorganisms, and the importance of earthworms in environmental management and their use in organic waste management. A straightforward and integrative approach has been taken to reviewing the literature of this important area (over 1400 references are cited). An extensive appendix has also been added summarizing the toxicity to earthworms of a wide range of chemicals. This book will continue to be an invaluable text for students of agriculture, soil science, ecology and invertebrate zoology.

Trichodorids constitute a group of nematodes, together with tylenchs and longidorids, that are parasitic on plants. The trichodorids form the smallest group in a number of genera, species and also individuals in the soil. However, they are not at all devoid of interest on both economical and zoological grounds, and they are being studied with increasing attention by agronomists and taxonomists. Trichodorids are important in strictly agronomical research, as they are able to cause direct damage to the root system of various crops; they are the agent of the docking disorder' of the sugar beet, a severe disease in Europe. They are also able to transmit various viruses to a series of important crops. These viruses differ from those transmitted by the longidorids and also - as opposed to longidorids - they are not strictly associated with one or a restricted number of species. Trichodorids therefore constitute a potentially active group of direct and indirect predators. The present book has been written by one of the leading specialists on the group, who has produced an up to date review of data on the morpho-anatomy of all species of the group. Moreover, the book is not, as is so often the case, based merely on a literature survey, but on the author's personal observations. The descriptions are supported by detailed drawings and micrographs and in many instances new characters of taxonomic significance have been identified. Attention to detail and keen observation have enabled the author to provide a considered and authoritative opinion on the taxonomy and systematics of the Trichodoridae. Further, the evaluation and selection of specific characteristics clearly demonstrate the importanceof reliable identification of species when undertaking investigations on geographical distribution, ecology and other facets of the biology of individual species, particularly those of economic importance. It is inevitable that in such an actively investigated taxon as the Trichodoridae, there will be a continuing addition of new species which in turn will encourage further expression of taxonomic opinion. If that proves to be so, Dr. Decraemer's book will provide a well-established foundation for any new opinion and will remain a standard reference for Trichodoridae as well as a substantial contribution to the science of nematology. Audience: Research workers and students of agriculture, horticulture and zoology, plant pathologists and extension nematology service.

The Third International Symposium on Cladocera, papers from which make up this volume, covered recent findings on the behaviour, life history, population genetics, reproduction, chemical communication, predator-prey interactions, epibionts, taxonomy, phylogeny, palaeolimnology and biogeography of this animal group. The Cladocera occupy an intermediate position in lake ecosystems, both as plankton and benthic organisms. Their often high abundance and their function, as transfer organisms from algae and dead organic matter to macro-invertebrates and fish, make them one of the most important organisms to affect the biological processes in freshwater ecosystems. The Cladocera living among the plankton often have recognizable distribution patterns and migrations; their size and brief life cycles make them popular in laboratory experiments; their cyclic parthenogenesis makes them suitable for many aspects of population genetics; and they are present as microfossils in lake sediments. All these features confer a unique status on the Cladocera as model organisms in many aspects of modern systematics and ecology.

Biologists since Darwin have been intrigued and confounded by the complex issues involved in the evolution and ecology of the social behavior of insects. The self-sacrifice of sterile workers in ant colonies has been particularly difficult for evolutionary biologists to explain. In this important new book, Andrew Bourke and Nigel Franks not only present a detailed overview of the current state of scientific knowledge about social evolution in ants, but also show how studies on ants have contributed to an understanding of many fundamental topics in behavioral ecology and evolutionary biology.

One of the substantial contributions of "Social Evolution in Ants" is its clear explanation of kin selection theory and sex ratio theory and their applications to social evolution in insects. Working to dispel lingering skepticism about the validity of kin selection and, more broadly, of "selfish gene" theory, Bourke and Franks show how these ideas underpin the evolution of both cooperation and conflict within ant societies. In addition, using simple algebra, they provide detailed explanations of key mathematical models. Finally, the authors discuss two relatively little-known topics in ant social biology: life history strategy and mating systems.

This comprehensive, up-to-date, and well-referenced work will appeal to all researchers in social insect biology and to scholars and students in the fields of entomology, behavioral ecology, and evolution.

Floricultural crops all over the world are challenged by a number of insect and mite pests. The pest scenario is changing, and with climate change the instances of new pest incidences have become a more common problem. Like other crops, the intensive cultivation of commercial flowers has accentuated pest problems, as farmers tend to use more agricultural chemicals, which, in turn, increase the problems of pesticide resistance, pest resurgence, and residues leading to health hazards. This volume, Advances in Pest Management in Commercial Flowers, looks at the major challenges and improvements in this growing area today. It first provides an informative overview of worldwide pests of important commercial flowers. It explores a number of important issues in this area, such as the role of climate change on insect pests of commercial flowers and the synthetic chemicals and their possible harmful effects on the environment.

During the past decade, the study of the chemical structures used by insects has advanced from a subject that could be reviewed in a single volume to a vastly more advanced level. This important new volume brings together a focused group of reviews that offer perspective on the most interesting advances in insect chemical ecology. Chemical Ecology of Insects 2 brings together an internationally respected group of experts covering such topics as chemoreception and integration, orientation mechanisms, plant-insect interactions and insect-insect interactions. An important benefit of these reviews lies in the identification of the boundaries of our current knowledge and the most profitable areas in which we should expect these areas to develop. This important work will appeal to entomologists and ecologists working directly with insects. In addition, plant scientists interested in the interaction of plants and insects will find much valuable information. The book is intended to benefit both field and laboratory researchers as well as advanced students.

This timely volume presents a comprehensive overview of the behavioral and physiological mechanisms underlying insect feeding. The single source for up-to-date information on the subject, it features contributions from the most authoritative scientists in the field worldwide. Taking an accessible approach focusing on the sequential stages of the feeding process, Regulatory Mechanisms in Insect Feeding explores the principles regulating feeding, and it offers information necessary for the development of more effective and environmentally safe methods to control the feeding of pest insects. The authors incorporate a spectrum of information in areas ranging from neurophysiology to insect control. An inclusive reference that reflects recent expansion and advances in the field, this book considers the problems encountered by insects when feeding on different types of food, as well as their adaptations for dealing with the problems introduces the cyclic nature of feeding activity and its components and provides the framework for the discussion of various control mechanisms discusses the underlying regulatory mechanisms of feeding in both the short and long terms in insects with a diversity of feeding habits synthesizes concepts and details of the basic mechanisms underlying insect feeding behavior that are essential for the development of pest management methods. Regulatory Mechanisms in Insect Feeding is the ideal text for graduate courses in entomology and zoology with a special focus on insect behavior, physiology, neurobiology, and control. It is also a valuable resource for ecologists interested in insect-plant and insect-animal interactions, for scientists concerned with the mechanismsbehind the physiology and behavior of feeding, and for those working in crop protection and pest management.

During the past decade, the study of the chemical structures used by insects has advanced from a subject that could be reviewed in a single volume to a vastly more advanced level. This important new volume brings together a focused group of reviews that offer perspective on the most interesting advances in insect chemical ecology. Chemical Ecology of Insects 2 brings together an internationally respected group of experts covering such topics as chemoreception and integration, orientation mechanisms, plant-insect interactions and insect-insect interactions. An important benefit of these reviews lies in the identification of the boundaries of our current knowledge and the most profitable areas in which we should expect these areas to develop. This important work will appeal to entomologists and ecologists working directly with insects. In addition, plant scientists interested in the interaction of plants and insects will find much valuable information. The book is intended to benefit both field and laboratory researchers as well as advanced students.

Ecology and Morphology of Copepods is organized under the following general topics: Behavior, Feeding, Genetics, Horizontal Variations, Morphology, Phylogeny, Reproduction, Seasonal Changes, Vertical Distribution, plus two special sessions on copepods of the genus Acartia and cyclopid/mosquito interactions. The Maxilliped Lecture, given by Dr. Arthur G. Humes of Boston University, clearly established the importance of copepods to the earth's organic diversity. The book consists of selected research articles from the different sections. The articles published here reflect the diverse research interests of copepodologists today, and are distinguished by their high quality. Their impact will ensure that this volume is consulted by a wide range of research biologists.

The dipteran family Chironomidae is the most widely distributed and frequently the most abundant group of insects in freshwater, with rep resentatives in both terrestrial and marine environments. A very wide range of gradients of temperature, pH, oxygen concentration, salinity, current velocity, depth, productivity, altitude and latitude have been exploited, by at least some chironomid species, and in grossly polluted environments chironomids may be the only insects present. The ability to exist in such a wide range of conditions has been achieved largely by behavioural and physiological adaptations with relatively slight morphological changes. It has been estimated that the number of species world-wide may be as high as 15000. This high species diversity has been attributed to the antiquity of the family, relatively low vagility leading to isolation, and evolutionary plasticity. In many aquatic ecosystems the number of chironomid species present may account for at least 50% of the total macroinvertebrate species recorded. This species richness, wide distribution and tolerance to adverse conditions has meant that the group is frequently recorded in ecological studies but taxonomic difficulties have in the past prevented non-specialist identification beyond family or subfamily level. Recent works, including genetic studies, have meant that the family is receiving much more attention globally."

In this volume outstanding specialists review the state of the art in nervous system research for all main invertebrate groups. They provide a comprehensive up-to-date analysis important for everyone working on neuronal aspects of single groups, as well as taking into account the phylogenesis of invertebrates. The articles report on recently gained knowledge about diversification in the invertebrate nervous systems, and demonstrate the analytical power of a comparative approach. Novel techniques in molecular and developmental biology are creating new perspectives that point toward a theoretical foundation for a modern organismic biology. The comparative approach, as documented here, will engage the interest of anyone challenged by the problem of structural diversification in biology.

Recent studies have shown that genetic polymorphisms play an important role in structuring the seasonal life cycles of insects, complementing an earlier emphasis on the effects of environmental factors. This book presents current ideas and recent research on insect life--cycle polymorphism in a series of carefully prepared chapters by international experts, covering the full breadth of the subject in order to give an up-to-date view of how life cycles are controlled and how they evolve. By consolidating our view of insect life--cycle polymorphism in this way, the book provides a staging point for further enquiries. The volume will be of interest to a wide variety of entomologists and other biologists interested in the control and evolution of life cycles and in understanding the extraordinarily complex ecological strategies of insects and other organisms.

This book was conceived to mark the Silver Jubilee of the British Butterfly Conservation Society. Interest in the conservation of butterflies has increased so rapidly that it is difficult to relate to the situation 25 years ago. Butterflies were on the decline in Britain, Europe and elsewhere but we lacked data on the extent of the decline and the underlying reasons, leaving us unable to implement effective conservation measures. An early recognition of the plight of British butterflies and moths led to the foundation of the society by a small group of conservationists in 1968. Today the society has over 10000 members, owns a number of reserves and sponsors research, conservation and monitoring activities at the local and national level. As part of the Silver Jubilee celebrations an international symposium was held at Keele University in September 1993 entitled 'Ecology and Conservation of Butterflies'. This symposium clearly showed how much important work has been done in recent years and also gave me the impression that the subject had reached a watershed. This was not because the decline of butterflies has stopped or even slowed down, far from it, the threat to our butterflies continues to increase from habitat destruction and intensification of land use. The watershed is in our understanding of the relationship between butterflies and their habitat.

This book is an identification guide to the arthropods (insects, mites, ticks, etc.) which affect the health of people and their domestic animals. It is designed for practical use on the laboratory bench and in the field. Coverage of organisms is world-wide, allowing the student to become familiar with and identify to genus level, all types of medical and veterinary pests. The book's easy to use format makes it useful also to the non-specialist and as a supplement to the conventional textbooks on medical and veterinary entomology.

This book will give an overview of insect ovaries, showing the diversities and the common traits in egg growth processes. The idea to write this book developed while looking at the flood of information which appeared in the early 1980s on early pattern formation in Drosophila embryos. At this time a significant breakthrough was made in studies of this little fly, combining molecular biological methods with classical and molecular genetics. The answers to questions about early pattern formation raised new questions about the architecture of ovaries and the growth of eggs within these ovaries. However, by concentrating only on Drosophila it is not possible to form an adequate picture of what is going on in insect ovaries, since the enormous diversity found among insects is not considered sufficiently. Almost forgotten, but the first to study the architecture of ovaries, was Alexander Brandt writing in 1878 in aber das Ei und seine Bildungsstaette (On the egg and its organ of development). More than 100 years later, a series of ten books or more would be required to survey all the serious informa tion we have today on insect oogenesis. Thus, this book is a personal selection and personal view on the theme, and the authors must be excused by all those scientists whose papers could not be included. The book briefly describes the ectodemes, i. e."

Phil Lester’s first book, The Vulgar Wasp, was about one of the world’s most hated insects. His second is about just the opposite - the honey bee, arguably one of our best-loved six-legged creatures. People have revered honey bees for centuries. Today we celebrate them with toys, postage stamps and campaigns to raise awareness; we dress up in large bee suits to protest the use of pesticides; we’ve even sent bees into space and watched as they adjusted to microgravity. Bees are one of the world’s most efficient pollinators. Their work is vital to the success of many food crops, and hence to the world’s economy. So we need to take seriously any threats to their health - including parasites, pathogens, predators and pesticides - and, guided by evidence at every turn, find a way to minimise harm and keep bees thriving. As Healthy Bee, Sick Bee shows, this is no small task.In this book, entomologist Phil Lester explores the wonderfully complex and sometimes brutally efficient life history of honey bees, and the problems they face in New Zealand and around the globe. What causes a beehive to collapse? Are pesticides as big a problem as they appear? What can we do to improve the health?

For more than 20 years insect/plant relations have been a focus for studies in ecology and evolution. The importance of insects as crop pests, and the great potential of insects for the biological control of weeds, have provided further impetus for work in this area. All this attention has resulted in books on various aspects of the topic, and reviews and research papers are abundant. So why write another book? It seems to us that, in the midst of all this activity, behavior has been neglected. We do not mean to suggest that there have not been admirable papers on behavior. The fact that we can write this book attests to that. But we feel that, too often, behavior is relegated to a back seat. In comparison to the major ecological and evolutionary questions, it may seem trivial. Yet the whole process of host-plant selection and host-plant specificity amongst insects depends on behavior, and selection for behavioral differences must be a prime factor in the evolution of host-plant specificity. In writing this book, we hope to draw attention to this central role of behavior and, hopefully, encourage a few students to attack some of the very difficult questions that remain unanswered.

This is the first book to provide a detailed treatment of the field of larval ecology. The 13 chapters use state-of-the-art reviews and critiques of nearly all of the major topics in this diverse and rapidly growing field. Topics include:
o patterns of larval diversity
o reproductive energetics
o spawning ecology
o life history theory
o larval feeding and nutrition
o larval mortality
o behavior and locomotion
o larval transport
o dispersal
o population genetics
o recruitment dynamics
o larval evolution
Written by the leading new scientists in the field, chapters define the current state of larval ecology and outline the important questions for future research.

Insects are the major component of the world's biodiversity. By their vast numbers of both species and individuals, they are vital determinants of the terrestrial ecological processes. Quantitatively, insects are important pointers for the species-rich geographical areas. Qualitatively, they are also important, whether the subjects of conservation themselves or as tools for identifying biotic areas with high endemism. "Insect Conservation Biology" covers a wide range of topics from single species to landscape conservation, and from rare butterflies to the benefits-and-risks of biocontrol agents. The approach is both positive and realistic, with insects being discussed in the contexts of sustainable development, agroecology and monitoring environmental change. Ethical issues surrounding insects are also considered as well as preservation technology and restoration ecology. Conservation circles have given too little attention to the ecological significance of insects, while entomologists have been employed mostly to control a tiny minority of species of insect pests. The realms of conservationist and entomologist are brought together in this book. This book should be of interest to conservation managers/biologists, entomologists, ecologists, and environmental scientists interested in biological diversity.

Grasslands comprise more than a quarter of the Earth's land surface. In addition to supporting a wide range of vertebrates such as domestic livestock and a variety of games species, grassland is the natural habitat for a wide range of invertebrate species, and this book considers those which occur in grassland and their impact on soil fertility and herbage growth. It describes grassland as a habitat for invertebrates, the groups which occur there and their abudance. An extensive literature on grassland invertebrates scattered through numerous scientific journals and reports is drawn on in an attempt to develop an overview. In the opening chapter the major grassland types are considered and the features which influence the distribution and abudance of the invertebrates which inhabit them are discussed. Next the major taxonomic groups are reviewed in turn, with a brief account of their biology and ecology and of their ecosystem role. Some general features of grassland invertebrate communities are then described and teh factors which influence th epopulation densities of their constituent species are considered. Particular attentionm is given to the ways in which populations are influenced by management practices. The final and largest chapterdeals with the various ways in which invertebrates influence important grassland processes through ingestion of organic matter, interaction with injurious species is considered, with particular emphasis on the potential for achieving this through manipulating grassland management practices.

Planthoppers include some of the most devastating pests of major agricultural crops throughout the world. One species, the rice brown planthopper, is among the most economically important pests in Asia. In past decades, government policies encouraged the control of rice planthoppers with synthetic pesticides, a tactic which promoted insecticide resistance and often led to the pesticide-induced resurgence of pest populations. To deter planthopper outbreaks, a more ecologically sound management strategy is being implemented, one based on a thorough investigation of population dynamics, natural enemies, and the genetics of host plant and insecticide adaptation. In the natural habitats of North America and Europe, scientists have also used planthoppers as model organisms to test ecological and evolutionary theory. The consequence of these diverse studies is an extremely scattered literature on planthoppers that has never been synthesized from an ecological perspective. This volume summarizes what is known about planthopper ecology and biological control. It takes a theoretical approach yet is deeply concerned with the application of theory to the practical problems of pest management.

A great deal is now known about the functional organization, physiology, reproduction, and development of barnacles. For the first time, this book brings to bear all aspects of this knowledge on our interpretation of the dynamics of barnacle evolution relating them to the fossil history and biogeography of the group.

Many mites possess extremely intricate life styles in close association with plant and animal hosts. Their polymorphism has made classification a challenge, and their ability to reproduce both sexually and asexually has made efforts to control their populations difficult. This, however, has given rise to theories to explain the origin and function of sexual reproduction in general. In numbers of species and geographic distribution, mites may even surpass the insects. In soils, they are a major component in the system for cycling nutrients. Unlike insects, they have invaded the marine environment. These and a number of other topics are explored in Mites. Because of their extremely small size, mites have been ignored during the development of major evolutionary and ecological theories. Yet mites routinely violate fundamental concepts such as heterochrony, sexual selection, the evolution of sex ratio, and ontogeny. Recent research methodologies have made it practical for the first time to perform experimental work with mites, and since they offer short generation times and rapid research results, they are excellent model systems. Mites announces these results and should appeal to professionals in entomology, acarology, ecology, population genetics, and evolutionary biology.

Surprising though it seems, the world faces almost as great a threat today from arthropod-borne diseases as it did in the heady days of the 1950s when global eradication of such diseases by eliminating their vectors with synthetic insecticides, particularly DDT, seemed a real possibility. Malaria, for example, still causes tremendous morbidity and mortality throughout the world, especially in Africa. Knowledge of the biology of insect and arachnid disease vectors is arguably more important now than it has ever been. Biological research directed at the development of better methods of control becomes even more important in the light of the partial failure of many control schemes that are based on insecticide- although not all is gloom, since basic biological studies have contributed enormously to the outstanding success of international control programmes such as the vast Onchocerciasis Control Programme in West Africa. It is a sine qua non for proper understanding of the epidemiology and successful vector control of any human disease transmitted by an arthropod that all concerned with the problem - medical entomologist, parasitologist, field technician - have a good basic understanding of the arthropod's biology. Knowledge will be needed not only of its direct relationship to any parasite or pathogen that it transmits but also of its structure, its life history and its behaviour - in short, its natural history. Above all, it will be necessary to be sure that it is correctly identified.

Darwin first raised the question of sex ratio evolution, and saw it as both important and enigmatic. He was, however, unable to make much headway with the problem and declared it a puzzle for the future. This particular future arrived about 60 years later, when R. A. Fisher (1930) pointed out that under autosomal inheritance half of the genes passed to zygotes in any generation come from males and half come from females. Fisher noted that this one-mother/one-father symmetry generates frequency dependent natural selection on sex ratio, resulting in an evolutionary equi librium in which half of the reproductive resources are devoted to daughters and half to sons. Although widespread interest in sex ratio as a phenotypic trait did not occur for another 30 years, it is difficult for us to overestimate the impor tance of Fisher's brief and characteristically cryptic remarks. Almost all of the innovations in thinking about sex ratio can be viewed as alterations of one or more implicit assumptions in Fisher's scheme. The present book on insect sex ratios is testimony to the fruitfulness of his original ideas and of their descendants."