The acridoid grasshoppers in general and the various species of swarm-forming locusts in particular have been among the most favoured subjects of insect physi ology, behaviour and ecology for many years. Several factors contribute to this popularity: their abundance in nature and their ease of culture in the laboratory, their relatively large size, and most of all, perhaps, their agricultural importance and the consequent intermittent availability of funds for their study. These factors together have inspired a large amount of experimental work, and this in tum has produced a new and often over-riding reason for working on acridoids -the huge body of available background information and know-how that has built up about these insects. This state of affairs is well seen in insect neurobiology. Only a restricted number of insect types are commonly used in this discipline, and originally most of them were selected for reasons of convenience and availability: grasshoppers, cock roaches, crickets, flies, bees and moths are the most important. Each of these in sects is the subject of the attentions of one or more major groups of neurobiolo gists, but neurobiological articles on acridoids probably exceed in number those on all other insects combined, at least if articles on the molecular biology of the nervous system of Drosophila are excluded.

The Biological Sciences are in the midst of a scientific rev olution. During the past decade under the rubric of molecu lar biology, chemistry and physics have assumed an integral role in biological research. This is especially true in ge netics, where the cloning of genes and the manipulation of genomic DNA have become in many organisms routine laboratory procedures. These noteworthy advances, it must be empha sized, especially in molecular genetics, are not autonomous. Rather, they have been accomplished with those organisms whose formal genetics has been documented in great detail. For the beginning student or the established investigator who is interested in pursuing eukaryote molecular genetic re search, Drosophila melanogaster, with its rich body of formal genetic information is one organism of choice. The book "Drosophila Genetics. A Practical Course" is an indispens able source of information for the beginner in the biology and formal genetics of Drosophila melanogaster. The scope of this guide, a revision and enlargement of the original German language version, is broad and instructive. The information included ranges from the simple, but necessary, details on how to culture and manipulate Drosophila flies to a series of more sophisticated genetic experiments. After completing the experiments detailed in the text, all students - neophyte or experienced - will be richly rewarded by having acquired a broad base of classical genetics information relevant for the biologist in its own right and prerequisite to Drosophila genetics research - formal and/or molecular. Davis, California, Melvin M."

Darwin first raised the question of sex ratio evolution, and saw it as both important and enigmatic. He was, however, unable to make much headway with the problem and declared it a puzzle for the future. This particular future arrived about 60 years later, when R. A. Fisher (1930) pointed out that under autosomal inheritance half of the genes passed to zygotes in any generation come from males and half come from females. Fisher noted that this one-mother/one-father symmetry generates frequency dependent natural selection on sex ratio, resulting in an evolutionary equi librium in which half of the reproductive resources are devoted to daughters and half to sons. Although widespread interest in sex ratio as a phenotypic trait did not occur for another 30 years, it is difficult for us to overestimate the impor tance of Fisher's brief and characteristically cryptic remarks. Almost all of the innovations in thinking about sex ratio can be viewed as alterations of one or more implicit assumptions in Fisher's scheme. The present book on insect sex ratios is testimony to the fruitfulness of his original ideas and of their descendants."

The increasing use of integrated crop management, often requiring a reduction in the reliance upon chemical control, means that the need to rapidly identify pest nematodes has never been greater. This second edition of this standard reference work familiar to all plant nematologists is therefore even more useful than its predecessor published in 1986. The in-depth description of the life histories of the genera of the Tylenchida have been retained and brought up-to-date through the inclusion of all the research carried out between the publication of the last edition and this new edition. This expanded edition includes detailed diagnoses of well over 200 genera and familial and ordinal groups and the book is heavily illustrated with drawings of type or representative species. These, together with comprehensive lists of species and genera and their synonymies provide the foundation for the status and validation of each taxon within the Tylenchida. A considerable amount of information is provided regarding the biology, ecology and pathogenicity of these parasites. This book is the only comprehensive reference work on this important group of nematodes.

This book presents the whole picture of the ecological and evolutionary study on the ground beetle group, the subgenus Ohomopterus of the genus Carabus, endemic to Japan. This flightless beetle group consists of many geographic races. They show divergence in key traits for reproductive isolation-body size and genital morphology, which leads to coexistence of two or more species. This beetle group provides an important material to study how a lineage of organisms diversify and form multi-species assemblage, and thereby multiply their species richness. The book introduces novel genomic approaches to resolve questions about evolution of Ohomopterus. The readers will find that this story of evolution in Carabus beetles revealed by recent approaches is much different from what was told in previous literature. Exploring different cases across a wide range of lineages is important in constructing a synthetic theory of species radiation and richness, including speciation and species coexistence. This study on Ohomopterus beetles contributes to the ongoing discussion to understand how and why species multiply and how species richness increases in one area of our planet.

Insect Chemical Ecology provides a comprehensive view of how natural selection acts upon interacting organisms and how particular physical and biological properties of chemical compounds act as constraints upon which natural selection may act. Individual chapters raise specific questions as to the nature of these interactions. The first part contains reviews on antagonistic and mutualistic chemical interactions, the raw materials' of chemical evolution, the economics of offensive and defensive chemicals, and neurobiology. The second part discusses particular problems such as the evolution of resistance, insect pollination, learning, pheromones, sequestration of semiochemicals, the role of microorganisms, sex attractants, the evolution of host races and biotypes, and the role of semiochemicals and the evolution of sociality of insects. The last chapter discusses the role of chemical-based pest management programs in an ecological and evolutionary framework.

In August 1988. the Sixth International Coral Reef Symposium was held in Townsville resulting in an influx of most of the world's coral reef sCientists to the city. We seized this opportunity at the Australian Institute of Marine Science to run a small workshop immediately before the symposium on the outbreaks of the crown-of-thorns starfish. Aeanthaster planei. We invited that small band of mathematicians who had been modelling the phenomenon, (and who may not have normally attended an international meeting so thoroughly dedicated to natural science) to meet with those SCientists who had been been actively working on the phenomenon in the field. John Casti notes in his delightful new book Alternate Realities (Wiley, 1989): 'If the natural role of the experimenter is to generate new observables by which we know the processes of Nature, and the natural role of the mathematician is to generate new formal structures by which we can represent these processes. then the system SCientist finds his niche by serving as a broker between the two. ' I think our book shows the fruits of that brokerage through the wide range of models explored within its pages. the high level of collaboration and interaction across disciplines evident in the individual papers, and in the emerging synthesis that reflects a far deeper understanding of this complex phenomenon than was possible even a few years ago.

Insect pest control continues to be a challenge for agricultural producers and researchers. Insect resistance to commonly used pesticides and the removal of toxic pesticides from the market have taken their toll on the ability of agricultural producers to produce high quality, pest-free crops within economical means. In addition to this, they must not endanger their workers or the environment. We depend on agriculture for food, feed, and fiber, making it an essential part of our economy. Many people take agriculture for granted while voicing concern over adverse effects of agricultural production practices on the environment. Insect Pest Management presents a balanced overview of environmentally safe and ecologically sound practices for managing insects. This book covers specific ecological measures, environmentally acceptable physical control measures, use of chemical pesticides, and a detailed account of agronomic and other cultural practices. It also includes a chapter on state-of-the-art integrated pest management based, a section on biological control, and lastly a section devoted to legal and legislative issues. Insect Pest Management approaches its subject in a systematic and comprehensive manner. It serves as a useful resource for professionals in the fields of entomology, agronomy, horticulture, ecology, and environmental sciences, as well as to agricultural producers, industrial chemists, and people concerned with regulatory and legislative issues.

This summary of the results of chromosomal analysis of various groups of parasitic wasps will be essential reading for those working in the field. It is the second edition of a monograph that was first published in Russian, and has been substantially updated with new data, conclusions and references.

The goal of this ground-breaking work is to document the author's study of the karyotype structure and chromosomal evolution of parasitic Hymenoptera. It also interprets the results obtained, from both routine and differential chromosome staining, for systematic and phylogenetic purposes.

Despite the huge importance of parasitic wasps, many problems of their taxonomy remain insufficiently studied. This is a situation that needs rectifying, as Hymenoptera are one of the largest and most taxonomically complicated groups of insects and they play a very important role in food chains as parasitoids of the overwhelming majority of insects and other arthropods. Parasitic wasps attack many pests of agriculture and forestry as well as certain beneficial insects. In addition, some secondarily phytophagous Hymenoptera damage certain cultivated plants.

The renowned entomologist and author of this volume, Vladimir Gokhman, has himself developed a particular technique for obtaining chromosomal preparations from adult parasitic wasps. This allowed the inclusion of a wide range of those insects into the study.

The karyotypic features of the majority of superfamilies and families as well as many lower taxa of parasitic Hymenoptera are listed, while the plesiomorphic and apomorphic character states of chromosome sets of parasitic wasps are revealed.

The structural features of karyotypes used for identifying and discriminating sibling species and similar forms are shown, as are the data on chromosome sets of about 400 species of parasitic Hymenoptera (including more than 200 micrographs). Also included is a comprehensive phylogenetic and taxonomic analysis of karyotypic information.

This book is unique because it demonstrates a range of karyotypic data for an insect group that seemed unattainable in this respect not so long ago. Rich in new facts, techniques and ideas, it will be of use to all those interested in karyology and Hymenoptera.

Honey Bees: Estimating the Environmental Impact of Chemicals is an updated account of the different strategies for assessing the ecotoxicity of xenobiotics against these social insects, which play a key role in both ecology and agriculture. In addition to the classical acute laboratory test, semi-field cage tests and full field funnel tests, new tests based mainly on behavioral responses are for the first time clearly described. Information on the direct and indirect effects on honey bees of radionuclides, heavy metals, pesticides, semi-volatile organic compounds and genetically modified plants is also presented.

Lucid line drawings and photos, 16 in fine color. Based on some 15 years of study and collection in both polar regions and most areas of the tropics and temperate zones. Detailed treatment is given on all known pelagic snails (aoubt 140 species) including: external anatomy, swimming and buoyancy mec

This major reference work contains essential information on arthropod-borne infections affecting humans and domesticated animals. The encyclopedia is a key reference source for anyone working in medical and veterinary science, and related fields. Features of "iThe Encyclopedia of Arthropod-transmitted Infections" are 150 entries, describing arboviral, viral, bacterial and rickettsial, spirochaetal, protozoal and filarial infections, and the vectors that transmit themInformation on disease distribution, clinical symptoms, diagnosis, transmission cycles, vector life-cycles, and treatment and control measures. Figures, tables and photographs illustrate the text. Following each entry is a selected bibliography, to aid further reading on the topic. Over 80 different international authors, with expertise in medicine, veterinary science, parasitology, entomology, epidemiology, microbiology, and zoology have contributed to the encyclopedia

Thisisnotintendedtobeacomprehensivetextbookofentomology andarachnology, butratheraconcisesynthesisofcertainbasic informationrequiredfor BSc(Hons)andMSc(Entomology) examinations. Theapproachisprimarilyfunctional: forinstance, theskeletalandwaterproofingpropertiesofthearthropodcuticle arediscussed, butnotitsbiochemistry;andIhaveincludedonly thosepointswithwhichIbelievealladvancedstudentsofthesub. jectoughttobefamiliar. Someaspectsaretodayregardedas outdated;othersdonotappearinanycurrenttexts, butIhave includedthembecauseIconsiderthemtobeimportant. Innoway, therefore, shouldthisberegardedasabookofreference. Tobe frank, itconsistsofamassofoversimplificationsandunqualified generalizationswhichareintendedtoclarifythecomplexprinciples underlyingthem. OncetheseprincipleshavebeenthorougWy grasped, thereaderwillhaveacquiredasufficientlybroadapproach tobeabletogetthebestvaluefrommoreadvancedtreatises. MythanksareduetoDrs. JohnDalingwater, AndrewMilner, andespeciallyPaulHillyardfortheiradviceonpalaeontological matters;toProfessorsEinarBursellforpermissiontouseFig. 8, takenfromhis An Introduction to Insect Physiology(Academic Press), NeilF. HadleyandtheEditorsof American Scientistto reproduceFigs. 52,53, andtoF. SchallerforFig. 42fromachap terhewroteinGupta(ed. ) Arthropod Phylogeny(VanNostrand Reinhold). Finally, IwouldliketoexpressmygratitudetoRoy Abrahamsforredrawingthefiguresshowinginsectwingvenations andtoEileenBerghfortypingthemanuscript. Thebookwas completedduringtenureofaLeverhulmeEmeritusFellowship whichIampleasedtoacknowledge, whileProfessorN. A. Mitchison FRSkindlyprovidedmewithaccommodationinUniversityCol. lege, London. J. L. CWUDSLEY-THOMPSON Contents 1 PalaeontologyandPhylogeny. . . . . . . . . . . . . . . . . . 1 1. 1 TheEarliestArthropodanFossils. . . . . . . . . . . . . . . . 1 1. 2 TheFirstTerrestrialArthropods. . . . . . . . . . . . . . . . . 2 1. 3 EvolutionintheArthropods. . . . . . . . . . . . . . . . . . . 4 1. 3. 1 EmbryologicalConsiderations. . . . . . . . . . . . . . . . . . 4 1. 3. 2 ComparativeMorphology. . . . . . . . . . . . . . . . . . . . . 5 1. 3. 3 MonophyleticorPolyphyleticOrigins. . . . . . . . . . . . . 6 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 2 ImplicationsofLiveonLand. . . . . . . . . . . . . . . 10 2. 1 TheSignificanceofSize 10 2. 1. 1 Size, SkeletonsandAllometry 11 2. 1. 2 AllometricGrowth 12 2. 2 WaterRelations 12 2. 3 TheConquestoftheLand 14 2. 4 . TheIntegument 14 2. 4. 1 TheEndocuticle 15 2. 4. 2 TheExocuticle 15 2. 4. 3 TheEpicuticle 16 2. 5 GrowthandEcdysis 18 2. 6 Respiration: Lung-BooksandTracheae 19 2. 7 NutritionandExcretion 21 2. 7. 1 Nutrition 21 2. 7. 2 Excretion 22 2. 8 EcologicalConsiderationsofSize 23 FurtherReading 25 3 TheConquestoftheLandbyCrustacea. . . . . . . . . . . . 26 3. 1 TypesofAdaptation. . . . 26 3. 2 TransitionfromWatertoLandinAmphipoda 26 3. 3 TransitionfromWatertoLandinDecapoda 28 3. 4 TransitionfromWatertoLandinIsopoda 30 3. 4. 1 Morphology 30 3. 4. 2Physiology. . . . . . . . . . . . . . . . . . . . . . . . . . . 31 3. 4. 3 Behaviour 34 VIII Contents 3. 5 Conclusion 35 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . 36 4 InsectPhylogenyandtheOriginofFlight. . . . 37 4. 1 AncestryofInsects 37 4. 2 TheOriginofWings 38 4. 2. 1 Apte10taandtheAncestryofSpiders 39 4. 3 Paranota1Theory 40 4. 4 TrachealGillTheory . 42 4. 5 SelectionforFlight 43 4. 6 PhylogenyoftheLowerInsectOrders . 44 4. 6. 1 Fossil

The present edition may be regarded as a descendant, much changed and greatly enlarged, of the late Dr A. D. Imms' Outlines of Entomology, first published in 1942. This went through three further editions without much change, but after the death of the original author a fifth, revised edition by Professor O. W. Richards and myself appeared in 1959 and a sixth in 1978. The book now appears in a considerably extended version in which I have tried to provide a more balanced introduction to the whole field of modern entomology by dealing with several aspects of the subject not discussed at all in previous editions. Thus, in addition to innumerable lesser changes in the sections on insect structure, function, development, classification and phylogeny, I have completely recast the earlier chapter on some important modes of life in insects. This now includes a far wider range of biological topics well exemplified by the insects and should, I hope, appeal not only to those already dedicated to entomology but also to others with more general biological interests. A completely new chapter on the biology of insect populations has also been added and may serve to indicate the debt which modern ecological theory owes to work on insect populations. It should hardly be necessary to apologize for introducing a certain amount of elementary mathematics into this account of a subject which is now among the most highly quantitative of biological disciplines.

When [ teach my undergraduate course in these animals belong together, and we nowlearn that Crustacea at the University of Copenhagen [always they may even encompass the speciose insects. With or tell my students: " Now, if on reading through the without insects the Crustacea are therefore unquestion- chapter on Crustacea you wonder, what could pos- ably one of the most important animal groups. Is this sibly be said in common about these animals, you why we study them? I think not. Our fascination with havegot it exactly right! Indeed, there is next to noth- Crustacea is even deeper rooted and not easily ana- ing that unite them". Crustaceans range in size from lyzed. But one thing is important to me in this context. much below millimeter size to the giant king crabs; As Ibrowse through aU the papers in this book it brings in morphology from the almost stereotyped segment up the memoryofthe highlyrewarding Summer Meet- repetition ofthe Remipedia to animals with a highly The Crustacean Society held in 2000 in Puerto ing of diverse array of appendages such as the Decapoda; Vallarta, Mexico, where I had the privilege to serve as in life form from suspension feeding anostracans to its president.

Palms constitute one of the largest botanical families, and include some of the world s most important economic plants. They are also unequalled as outdoor and indoor ornamental plants, and include many species that are essential components of the ecosystems of tropical and other warm regions. This book reviews the inter-relationships between palms and insects, emphasizing the similarities in different world regions. The host plants, distribution, and bionomics of representative insects are discussed according to their feeding sites on palms (foliage, flowers, fruits, and stems) and their taxonomic groups. Host and distribution records for the most extensively represented insect families on palms are tabulated. Pest management and field techniques are also covered. This book is recommended reading for tropical biologists and agriculturalists, including entomologists, horticulturists and tropical ecologists as well as palm nursery growers, managers and enthusiasts."

The authors are two of the best taxonomic experts in crustacean decapods, who have established the actual taxonomy and systematic of numerous and important groups of decapods. They have introduced new views and approaches on the taxonomy, phylogeny and biogeography of these crustaceans. This international edition adds diagnoses and identification keys to all world families and genera: this information simply doesn't exist in any other book. During the last decade, the world register of marine species (WORMS) has created the list of known species and promoted the publication of numerous compilations on crustacean decapods. However, except specialists of each taxa, very few colleagues can identify correctly the new families and genera - there are no keys to these new groups. This book is the first to contain these world keys and is therefore a life rope for students, marine biologists, taxonomists and naturalists. The mixture of taxonomic line drawings, color illustrations and images of real-life specimens is extremely helpful for identification, especially when working in the field.

The nervous system is particularly fascinating for many biologists because it controls animal characteristics such as movement, behavior, and coordinated thinking. Invertebrate neurobiology has traditionally been studied in specific model organisms, whilst knowledge of the broad diversity of nervous system architecture and its evolution among metazoan animals has received less attention. This is the first major reference work in the field for 50 years, bringing together many leading evolutionary neurobiologists to review the most recent research on the structure of invertebrate nervous systems and provide a comprehensive and authoritative overview for a new generation of researchers. Presented in full colour throughout, Structure and Evolution of Invertebrate Nervous Systems synthesizes and illustrates the numerous new findings that have been made possible with light and electron microscopy. These include the recent introduction of new molecular and optical techniques such as immunohistochemical staining of neuron-specific antigens and fluorescence in-situ-hybridization, combined with visualization by confocal laser scanning microscopy. New approaches to analysing the structure of the nervous system are also included such as micro-computational tomography, cryo-soft X-ray tomography, and various 3-D visualization techniques. The book follows a systematic and phylogenetic structure, covering a broad range of taxa, interspersed with chapters focusing on selected topics in nervous system functioning which are presented as research highlights and perspectives. This comprehensive reference work will be an essential companion for graduate students and researchers alike in the fields of metazoan neurobiology, morphology, zoology, phylogeny and evolution.

This balanced, comprehensive account traces the alterations in body form that insects undergo as they adapt to seasonal change, exploring both theoretical aspects and practical issues, such as the impact of seasonality on insect pest management.

Beetles, the most diverse group of insects, are often abundant in terrestrial and freshwater ecosystems. Many species are under threat from human changes to natural environments, and some are valuable tools in conservation, because they respond rapidly to changes that occur. Knowledge of these responses, of both abundance and composition of assemblages, enable use of some beetles to monitor environmental changes. Beetles impinge on humanity on many ways: as cultural objects, desirable collectables, major pests and competitors for resources need by people, as beneficial consumers of other pests, and by ensuring the continuity of vital ecological processes. This book is the first major global overview of the importance of conservation of beetles, and brings together much hitherto scattered information to demonstrate the needs for conservation, and how it may be approached. It is a source of value to students, research workers, conservation biologists and ecosystem managers as an introduction to the richness and importance of this predominant component of invertebrate life.

This book attempts to summarize what we know about inseet-plant relationships without beeoming too involved with untestable hypo- theses. It is not intended to be eomprehensive and we have deliberately excluded diseussion of aquatie organisms and fungi. Our definition of inseet herbivores is intentionally broad. It includes all inseets whieh feed on plants, although we have emphasized those whieh feed primarily on the photosynthetie tissues. Some referenee is made to seed predation but pollination eeology is excluded. We thank Ors P.H. Smith and M. Luxton for their helpful eommenb on the manuseript but we accept full responsibility for any mistakes whieh may remain. Finally, we thank the various publishers and a uthor~ who gave us permission to use copyright material. 7 I Introduction The net primary produetion of the 300000 speeies of vaseular plant whieh inhabit the dry land surfaee of the earth has been estimated at 9 about 115 x 10 t per annum. This represents a massive resouree potentially available for exploitation by the herbivorous inseets, which themselves probably number in exeess of 500000 species.

Cockroaches are ideal subjects for laboratory investigation at all educational levels. Compared with many other laboratory animals, cockroaches are easily and inexpensively maintained and cultured and require relatively little space. They are hardy and are readily available. The purpose of this book is to provide background material and experimental leads for utilizing cockroaches in the teaching laboratory and in designing research projects. The level of difficulty of the experiments varies according to the depth of understanding desi red by the instructor. In most cases at least a part of each experiment or technique can be incorporated into the laboratory component of elementary, high school or college curriculum. Sections of the lab book are appropriate for courses in Animal Behavior, Entomology, Organismic Biology and Insect Physiology. Aside from this main purpose, the book also provides a wealth of experimental ideas and techniques for a scientist at any level of education. Lawrence, Kansas June 15, 1981 W. J. B. ACKNOWLEDGEMENTS. Virtually all graduate students who have worked on cockroach research in my laboratory have knowingly or unknowingly contributed to this book. The most important contribution was from Sandy Jones McPeak, who encouraged me to finish the project. Segments of various chapters were conceived, developed or reviewed by Michael D. Breed, Sandy Jones McPeak, Michael K. Rust, Coby Schal, Thomas R. Tobin, W. Alexander Hawkins, Gary R. Sams and Chris Parsons Sams.

The invasive Harlequin ladybird is an eye-catching and beautiful species, but it can be very difficult to identify, with huge variation in colouration and pattern. This comprehensive photographic field guide is the first complete guide to identifying Harlequin ladybirds found in Britain and Ireland. It also covers all the other 25 conspicuous ladybird species that occur. Detailed, informative and beautifully illustrated with more than 350 remarkable colour photographs including ladybird developmental time sequences and numerous images of each ladybird species Ground-breaking quick key for Harlequin identification Includes a photographic guide to the larvae of all 26 species - the first of its kind Easy to follow comprehensive descriptions and explanations of ladybird lifecycles and habits This clear, user-friendly field guide is ideal for anyone interested in learning how to identify a Harlequin ladybird, from beginners to more experienced entomologists, students and professionals. This book is ideal for learning to recognise and understand this charismatic and significant beetle.

This volume provides an abundance of valuable first-hand information about the diversity, biology, ecology culture of the portunid crabs of the word seas. Marine crabs play an important role directly or indirectly in the livelihood of millions of people around the world. They have been reported to make up about 20% of all marine crustaceans caught, farmed, and consumed worldwide. Among these marine crabs, portunid crabs (or swimming crabs of the family Portunidae) assume greater significance in the marine industry, owing to their delicate meat with nutritional qualities. Although several species of portunid crabs are edible and commercially important, only a few species have been widely cultivated. This is largely due to the lack of information on the biology of portunid crabs. Keeping this in view, this volume presents the biology and aquaculture of marine portunid crabs. This volume will be of great use for researchers and students of disciplines such as fisheries science, marine biology, aquatic biology and fisheries, and zoology, and will also serve as a standard reference for college, university, and research libraries around the world.