This major reference work contains essential information on arthropod-borne infections affecting humans and domesticated animals. The encyclopedia is a key reference source for anyone working in medical and veterinary science, and related fields. Features of "iThe Encyclopedia of Arthropod-transmitted Infections" are 150 entries, describing arboviral, viral, bacterial and rickettsial, spirochaetal, protozoal and filarial infections, and the vectors that transmit themInformation on disease distribution, clinical symptoms, diagnosis, transmission cycles, vector life-cycles, and treatment and control measures. Figures, tables and photographs illustrate the text. Following each entry is a selected bibliography, to aid further reading on the topic. Over 80 different international authors, with expertise in medicine, veterinary science, parasitology, entomology, epidemiology, microbiology, and zoology have contributed to the encyclopedia

Lucid line drawings and photos, 16 in fine color. Based on some 15 years of study and collection in both polar regions and most areas of the tropics and temperate zones. Detailed treatment is given on all known pelagic snails (aoubt 140 species) including: external anatomy, swimming and buoyancy mec

The present edition may be regarded as a descendant, much changed and greatly enlarged, of the late Dr A. D. Imms' Outlines of Entomology, first published in 1942. This went through three further editions without much change, but after the death of the original author a fifth, revised edition by Professor O. W. Richards and myself appeared in 1959 and a sixth in 1978. The book now appears in a considerably extended version in which I have tried to provide a more balanced introduction to the whole field of modern entomology by dealing with several aspects of the subject not discussed at all in previous editions. Thus, in addition to innumerable lesser changes in the sections on insect structure, function, development, classification and phylogeny, I have completely recast the earlier chapter on some important modes of life in insects. This now includes a far wider range of biological topics well exemplified by the insects and should, I hope, appeal not only to those already dedicated to entomology but also to others with more general biological interests. A completely new chapter on the biology of insect populations has also been added and may serve to indicate the debt which modern ecological theory owes to work on insect populations. It should hardly be necessary to apologize for introducing a certain amount of elementary mathematics into this account of a subject which is now among the most highly quantitative of biological disciplines.

Thisisnotintendedtobeacomprehensivetextbookofentomology andarachnology, butratheraconcisesynthesisofcertainbasic informationrequiredfor BSc(Hons)andMSc(Entomology) examinations. Theapproachisprimarilyfunctional: forinstance, theskeletalandwaterproofingpropertiesofthearthropodcuticle arediscussed, butnotitsbiochemistry;andIhaveincludedonly thosepointswithwhichIbelievealladvancedstudentsofthesub. jectoughttobefamiliar. Someaspectsaretodayregardedas outdated;othersdonotappearinanycurrenttexts, butIhave includedthembecauseIconsiderthemtobeimportant. Innoway, therefore, shouldthisberegardedasabookofreference. Tobe frank, itconsistsofamassofoversimplificationsandunqualified generalizationswhichareintendedtoclarifythecomplexprinciples underlyingthem. OncetheseprincipleshavebeenthorougWy grasped, thereaderwillhaveacquiredasufficientlybroadapproach tobeabletogetthebestvaluefrommoreadvancedtreatises. MythanksareduetoDrs. JohnDalingwater, AndrewMilner, andespeciallyPaulHillyardfortheiradviceonpalaeontological matters;toProfessorsEinarBursellforpermissiontouseFig. 8, takenfromhis An Introduction to Insect Physiology(Academic Press), NeilF. HadleyandtheEditorsof American Scientistto reproduceFigs. 52,53, andtoF. SchallerforFig. 42fromachap terhewroteinGupta(ed. ) Arthropod Phylogeny(VanNostrand Reinhold). Finally, IwouldliketoexpressmygratitudetoRoy Abrahamsforredrawingthefiguresshowinginsectwingvenations andtoEileenBerghfortypingthemanuscript. Thebookwas completedduringtenureofaLeverhulmeEmeritusFellowship whichIampleasedtoacknowledge, whileProfessorN. A. Mitchison FRSkindlyprovidedmewithaccommodationinUniversityCol. lege, London. J. L. CWUDSLEY-THOMPSON Contents 1 PalaeontologyandPhylogeny. . . . . . . . . . . . . . . . . . 1 1. 1 TheEarliestArthropodanFossils. . . . . . . . . . . . . . . . 1 1. 2 TheFirstTerrestrialArthropods. . . . . . . . . . . . . . . . . 2 1. 3 EvolutionintheArthropods. . . . . . . . . . . . . . . . . . . 4 1. 3. 1 EmbryologicalConsiderations. . . . . . . . . . . . . . . . . . 4 1. 3. 2 ComparativeMorphology. . . . . . . . . . . . . . . . . . . . . 5 1. 3. 3 MonophyleticorPolyphyleticOrigins. . . . . . . . . . . . . 6 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 2 ImplicationsofLiveonLand. . . . . . . . . . . . . . . 10 2. 1 TheSignificanceofSize 10 2. 1. 1 Size, SkeletonsandAllometry 11 2. 1. 2 AllometricGrowth 12 2. 2 WaterRelations 12 2. 3 TheConquestoftheLand 14 2. 4 . TheIntegument 14 2. 4. 1 TheEndocuticle 15 2. 4. 2 TheExocuticle 15 2. 4. 3 TheEpicuticle 16 2. 5 GrowthandEcdysis 18 2. 6 Respiration: Lung-BooksandTracheae 19 2. 7 NutritionandExcretion 21 2. 7. 1 Nutrition 21 2. 7. 2 Excretion 22 2. 8 EcologicalConsiderationsofSize 23 FurtherReading 25 3 TheConquestoftheLandbyCrustacea. . . . . . . . . . . . 26 3. 1 TypesofAdaptation. . . . 26 3. 2 TransitionfromWatertoLandinAmphipoda 26 3. 3 TransitionfromWatertoLandinDecapoda 28 3. 4 TransitionfromWatertoLandinIsopoda 30 3. 4. 1 Morphology 30 3. 4. 2Physiology. . . . . . . . . . . . . . . . . . . . . . . . . . . 31 3. 4. 3 Behaviour 34 VIII Contents 3. 5 Conclusion 35 FurtherReading. . . . . . . . . . . . . . . . . . . . . . . . 36 4 InsectPhylogenyandtheOriginofFlight. . . . 37 4. 1 AncestryofInsects 37 4. 2 TheOriginofWings 38 4. 2. 1 Apte10taandtheAncestryofSpiders 39 4. 3 Paranota1Theory 40 4. 4 TrachealGillTheory . 42 4. 5 SelectionforFlight 43 4. 6 PhylogenyoftheLowerInsectOrders . 44 4. 6. 1 Fossil

When [ teach my undergraduate course in these animals belong together, and we nowlearn that Crustacea at the University of Copenhagen [always they may even encompass the speciose insects. With or tell my students: " Now, if on reading through the without insects the Crustacea are therefore unquestion- chapter on Crustacea you wonder, what could pos- ably one of the most important animal groups. Is this sibly be said in common about these animals, you why we study them? I think not. Our fascination with havegot it exactly right! Indeed, there is next to noth- Crustacea is even deeper rooted and not easily ana- ing that unite them". Crustaceans range in size from lyzed. But one thing is important to me in this context. much below millimeter size to the giant king crabs; As Ibrowse through aU the papers in this book it brings in morphology from the almost stereotyped segment up the memoryofthe highlyrewarding Summer Meet- repetition ofthe Remipedia to animals with a highly The Crustacean Society held in 2000 in Puerto ing of diverse array of appendages such as the Decapoda; Vallarta, Mexico, where I had the privilege to serve as in life form from suspension feeding anostracans to its president.

This balanced, comprehensive account traces the alterations in body form that insects undergo as they adapt to seasonal change, exploring both theoretical aspects and practical issues, such as the impact of seasonality on insect pest management.

The nervous system is particularly fascinating for many biologists because it controls animal characteristics such as movement, behavior, and coordinated thinking. Invertebrate neurobiology has traditionally been studied in specific model organisms, whilst knowledge of the broad diversity of nervous system architecture and its evolution among metazoan animals has received less attention. This is the first major reference work in the field for 50 years, bringing together many leading evolutionary neurobiologists to review the most recent research on the structure of invertebrate nervous systems and provide a comprehensive and authoritative overview for a new generation of researchers. Presented in full colour throughout, Structure and Evolution of Invertebrate Nervous Systems synthesizes and illustrates the numerous new findings that have been made possible with light and electron microscopy. These include the recent introduction of new molecular and optical techniques such as immunohistochemical staining of neuron-specific antigens and fluorescence in-situ-hybridization, combined with visualization by confocal laser scanning microscopy. New approaches to analysing the structure of the nervous system are also included such as micro-computational tomography, cryo-soft X-ray tomography, and various 3-D visualization techniques. The book follows a systematic and phylogenetic structure, covering a broad range of taxa, interspersed with chapters focusing on selected topics in nervous system functioning which are presented as research highlights and perspectives. This comprehensive reference work will be an essential companion for graduate students and researchers alike in the fields of metazoan neurobiology, morphology, zoology, phylogeny and evolution.

Beetles, the most diverse group of insects, are often abundant in terrestrial and freshwater ecosystems. Many species are under threat from human changes to natural environments, and some are valuable tools in conservation, because they respond rapidly to changes that occur. Knowledge of these responses, of both abundance and composition of assemblages, enable use of some beetles to monitor environmental changes. Beetles impinge on humanity on many ways: as cultural objects, desirable collectables, major pests and competitors for resources need by people, as beneficial consumers of other pests, and by ensuring the continuity of vital ecological processes. This book is the first major global overview of the importance of conservation of beetles, and brings together much hitherto scattered information to demonstrate the needs for conservation, and how it may be approached. It is a source of value to students, research workers, conservation biologists and ecosystem managers as an introduction to the richness and importance of this predominant component of invertebrate life.

Butterflies of Maine and the Canadian Maritime Provinces introduces readers to over one hundred and twenty butterfly species documented in the Acadian region. Including contributions from researchers and community scientists, this volume is indispensable for anyone interested in the study and conservation of these ecologically important insects. This user-friendly guide features: • The first annotated checklist of the species and subspecies of Maine, New Brunswick, Nova Scotia, and Prince Edward Island • Species accounts covering habitat, behavior, threats, and more • Color photographs, flight histograms, and distribution maps

Learn to Identify Tiger Beetles with This Field Guide We are all very lucky that tiger beetles aren't the size of German Shepherds! These ferocious, pint-sized predators are beautifully marked, and some are extremely colorful. They are fascinating to watch and easy to find. Join in the fun with Tiger Beetles of Minnesota, Wisconsin & Michigan by Mathew Brust with full-color photography by Mike Reese. All 21 species found in Minnesota, Wisconsin, and Michigan are covered in detail, with loads of amazing natural history and the information you need to identify what you see. Inside You'll Find All 21 species of tiger beetles found in Minnesota, Wisconsin & Michigan Innovative format that makes field identification a snap Fieldmark arrows that point out the best distinguishing characteristics The best habitats to look for tiger beetles More than 180 color photos and detailed range maps for all species Elytra comparison chart for easy IDs "Tiger beetling" has become very popular with naturalists across the country. Get in on the fun with this handy field guide that conveniently fits in your backpack or back pocket.

A how-to book on an exhilarating outdoor activity and a unique meditation on the pleasures of the natural world Following the Wild Bees is a delightful foray into the pastime of bee hunting, an exhilarating outdoor activity that used to be practiced widely but which few people know about today. Weaving informative discussions of bee biology with colorful anecdotes, personal insights, and beautiful photos, Thomas Seeley describes the history and science behind this lost pastime and how anyone can do it. The bee hunter's reward is a thrilling encounter with nature that challenges mind and body while also giving insights into the remarkable behavior of honey bees living in the wild. Whether you're a bee enthusiast or just curious about the natural world, this book is the ideal companion for newcomers to bee hunting and a rare treat for armchair naturalists.

This book attempts to summarize what we know about inseet-plant relationships without beeoming too involved with untestable hypo- theses. It is not intended to be eomprehensive and we have deliberately excluded diseussion of aquatie organisms and fungi. Our definition of inseet herbivores is intentionally broad. It includes all inseets whieh feed on plants, although we have emphasized those whieh feed primarily on the photosynthetie tissues. Some referenee is made to seed predation but pollination eeology is excluded. We thank Ors P.H. Smith and M. Luxton for their helpful eommenb on the manuseript but we accept full responsibility for any mistakes whieh may remain. Finally, we thank the various publishers and a uthor~ who gave us permission to use copyright material. 7 I Introduction The net primary produetion of the 300000 speeies of vaseular plant whieh inhabit the dry land surfaee of the earth has been estimated at 9 about 115 x 10 t per annum. This represents a massive resouree potentially available for exploitation by the herbivorous inseets, which themselves probably number in exeess of 500000 species.

Cockroaches are ideal subjects for laboratory investigation at all educational levels. Compared with many other laboratory animals, cockroaches are easily and inexpensively maintained and cultured and require relatively little space. They are hardy and are readily available. The purpose of this book is to provide background material and experimental leads for utilizing cockroaches in the teaching laboratory and in designing research projects. The level of difficulty of the experiments varies according to the depth of understanding desi red by the instructor. In most cases at least a part of each experiment or technique can be incorporated into the laboratory component of elementary, high school or college curriculum. Sections of the lab book are appropriate for courses in Animal Behavior, Entomology, Organismic Biology and Insect Physiology. Aside from this main purpose, the book also provides a wealth of experimental ideas and techniques for a scientist at any level of education. Lawrence, Kansas June 15, 1981 W. J. B. ACKNOWLEDGEMENTS. Virtually all graduate students who have worked on cockroach research in my laboratory have knowingly or unknowingly contributed to this book. The most important contribution was from Sandy Jones McPeak, who encouraged me to finish the project. Segments of various chapters were conceived, developed or reviewed by Michael D. Breed, Sandy Jones McPeak, Michael K. Rust, Coby Schal, Thomas R. Tobin, W. Alexander Hawkins, Gary R. Sams and Chris Parsons Sams.

The authors are two of the best taxonomic experts in crustacean decapods, who have established the actual taxonomy and systematic of numerous and important groups of decapods. They have introduced new views and approaches on the taxonomy, phylogeny and biogeography of these crustaceans. This international edition adds diagnoses and identification keys to all world families and genera: this information simply doesn't exist in any other book. During the last decade, the world register of marine species (WORMS) has created the list of known species and promoted the publication of numerous compilations on crustacean decapods. However, except specialists of each taxa, very few colleagues can identify correctly the new families and genera - there are no keys to these new groups. This book is the first to contain these world keys and is therefore a life rope for students, marine biologists, taxonomists and naturalists. The mixture of taxonomic line drawings, color illustrations and images of real-life specimens is extremely helpful for identification, especially when working in the field.

Courses on the invertebrates have two principal aims: (1) to introduce students to the diversity of animal life and (2) to make them aware that organisms are marvellously integrated systems with evolutionary pasts and ecological presents. This text is concerned exclusively with the second aim and assumes that the reader will already know something about the diversity and classification of invertebrates. Concepts of whole-organism function, metabolism and adaptation form the core of the subject-matter and this is also considered in an ecological setting. Hence, the approach is multi-disciplinary, drawing from principles normally restricted to comparative morphology and physiology, ecology and evolutionary biology. Invertebrate courses, as with all others in a science curriculum, also have another aim - to make students aware of the general methods of science. And these I take to be associated with the so-calledhypothetico deductive programme. Here, therefore, I make a conscious effort to formulate simple, some might say naive, hypotheses and to confront them with quantitative data from the real world. There are, for example, as many graphs in the book as illustrations of animals. My aim, though, has not been to test out the principles of Darwinism, but rather to sharpen our focus on physiological adaptations, given the assumption that Darwinism is approximately correct. Whether or not I succeed remains for the reader to decide."

This volume provides an abundance of valuable first-hand information about the diversity, biology, ecology culture of the portunid crabs of the word seas. Marine crabs play an important role directly or indirectly in the livelihood of millions of people around the world. They have been reported to make up about 20% of all marine crustaceans caught, farmed, and consumed worldwide. Among these marine crabs, portunid crabs (or swimming crabs of the family Portunidae) assume greater significance in the marine industry, owing to their delicate meat with nutritional qualities. Although several species of portunid crabs are edible and commercially important, only a few species have been widely cultivated. This is largely due to the lack of information on the biology of portunid crabs. Keeping this in view, this volume presents the biology and aquaculture of marine portunid crabs. This volume will be of great use for researchers and students of disciplines such as fisheries science, marine biology, aquatic biology and fisheries, and zoology, and will also serve as a standard reference for college, university, and research libraries around the world.

It was gratifying to be invited to prepare a third edition of this book, which first appeared in 1951. Preliminary discussions with the publishers, however, revealed a considerable challenge in the present high costs of printing, so that changes and some improvements were clearly necessary to justify the venture. It was immediately apparent that the chapter on chemical control measures would have to be substantially re-written, because of the great changes in usage due to resistance and the regulations introduced to prevent environmental pollution. Also, I decided to expand the scope of the book by increased coverage of the pests of continental Europe and North America, including some new figures and keys in the Appendix. These two undertakings resulted in considerable expansion in length, with about 370 new references and 250 additional specific names in the Index. In order to avoid too alarming an increase in price, I decided to sacrifice three chapters from the earlier editions: those dealing with the structure and classification of insects, their anatomy and physiology, and their ecology. Readers who require basic biological information on insects should buy one of the various short introductions to entomology available.

An analysis of the particular topics in biological control programmes all over the world indicates an increased interest in the utilization of aphid parasites. Besides the so-called traditional biological control of introduced aphids, there appears a situation which could perhaps be called a renaissance of biocontrol, i. e. the utilization of biotic agents against insecticide-resistant populations of aphid pests. The rapid increase in the amount of information as well as the necessity of syntheti cal papers are the well-known features in today's entomology. This requirement is much more topical in the groups where brief summarized information is needed for research workers in applied branches. The author has been well aware of all these difficulties and requirements owing to his own experience both in basic and applied research. Several years ago he decided to summarize our knowledge on the aphid parasites of the world by elaborating synthetical studies on the particular zoogeographical areas. These papers have been intended to represent annotated reviews of the parasite fauna, distribution, biologies and utilization in aphid pest management, with keys to genera and species, hos- parasite catalogue, and a list of references added. Naturally, these studies are only relatively updated, with respect to a certain deadline from which the research may be further continued. At present, the Far East Asian and the Mediterranean areas have been reviewed in this or at least in a similar manner.

The book summarizes the history of biological control in Latin America and the Caribbean. Few publications provide historical detail and the records are, therefore, fragmented until now. By bringing information together in this book, we offer a more complete picture of important developments in biological control on this continent. There are a wealth of text, tables and references about the history of such projects, and which were successful and which failed. This will help plan future biocontrol projects. An overview is provided of the current situation in biological control for many Latin American and Caribbean countries, revealing an astonishing level of practical biological control applied in the region, making it the largest area under biological control worldwide. The final part describes new developments and speculates about the future of biological control in Latin America and the Caribbean.

There has been a considerable upsurge in interest in insect bio chemistry and physiology in recent years and this has been reflected in a notable expansion in the number of original papers in this field. Whereas insect physiology has tended to receive ample attention from reviewers, the same has not always been true for the more biochemical aspects of insect research. This book is a venture to help redress the balance. No attempt has been made to cover all aspects of insect biochemistry, but rather a few topics have been selected which seemed to us to merit a review at the present time. One reason for this increased interest in insect biochemistry is perhaps the growing realization that insects can be very useful organisms to act as model systems for the experimental study of general biochemical principles. One remembers, for instance, that Keilin's perceptive observations on the flight muscles of living bees and wax moths led to his discovery of the cytochromes. The fundamental unity of biochemistry has long been accepted as a dogma by the faithful and the insect kingdom provides no exception to it. The main biochemical processes in insects are being revealed as essentially the same as in other life forms but, as so often found in comparative biochemistry, there are interesting variations on the central theme."

Winner of the Association of American Publishers PROSE Award for Excellence in Biological and Life Sciences A world-renowned researcher of animal behavior reveals the extraordinary orienteering skills of desert ants, offering a thrilling account of the sophisticated ways insects function in their natural environments. Cataglyphis desert ants are agile ultrarunners who can tolerate near-lethal temperatures when they forage in the hot midday sun. But it is their remarkable navigational abilities that make these ants so fascinating to study. Whether in the Sahara or its ecological equivalents in the Namib Desert and Australian Outback, the Cataglyphis navigators can set out foraging across vast expanses of desert terrain in search of prey, and then find the shortest way home. For almost half a century, Rudiger Wehner and his collaborators have devised elegant experiments to unmask how they do it. Through a lively and lucid narrative, Desert Navigator offers a firsthand look at the extraordinary navigational skills of these charismatic desert dwellers and the experiments that revealed how they strategize and solve complex problems. Wehner and his team discovered that these insect navigators use visual cues in the sky that humans are unable to see, the Earth's magnetic field, wind direction, a step counter, and panoramic "snapshots" of landmarks, among other resources. The ants combine all of this information to steer an optimal course. At any given time during their long journey, they know exactly where to go. It is no wonder these nimble and versatile creatures have become models in the study of animal navigation. Desert Navigator brings to light the marvelous capacity and complexity found in these remarkable insects and shows us how mini brains can solve mega tasks.

In his preface to early editions of this book, the late Dr. A. D. Imms said that he intended it to be an elementary account of entomology as a branch of general biology. He had especiaHy in mind the needs of university students of zoology and agriculture, as weH as those intending later to specialize in entomology, and he suggested that the book might also interest teachers of advanced biology in schools. These general aims and the balance between the different aspects of the subject have changed linIe in this and in our previous revision. We have, however, tried to bring the present edition up to date on the lines of our revised tenth edition of Imms' General Textbook 0/ Entomology, published in 1977. The text has been entirely re-set and eleven illustrations have been replaced by new figures. The same orders of insects are recognized as in the last edition, but the sequence in which the Endopterygote groups appear has been changed to reflect more accurately their probable evolutionary relationships. Many small changes and some addi tions have been made in the physiological sections, the chapter on the origin and phylogeny of insects has been rewritten, and a new bibliography provides a selection of modern references for the in tending specialist. It has been our object to make these alterations without materially increasing the length of the book or its level of difficulty."

seem as appropriate now as the original balance was when Dr A. D. Imms' textbook was first published over fifty years ago. There are 35 new figures, all based on published illustrations, the sources of which are acknowledged in the captions. We are grateful to the authors concerned and also to Miss K. Priest of Messrs Chapman & Hall, who saved us from many errors and omissions, and to Mrs R. G. Davies for substantial help in preparing the bibliographies and checking references. London O.W.R. May 1976 R.G.D. Part I ANATOMY AND PHYSIOLOGY Chapter I INTRODUCTION Definition of the Insecta (Hexapoda) The insects are tracheate arthropods in which the body is divided into head, thorax and abdomen. A single pair of antennae (homologous with the anten nules of the Crustacea) is present and the head also bears a pair of mandibles and two pairs of maxillae, the second pair fused medially to form the labium. The thorax carries three pairs of legs and usually one or two pairs of wings. The abdomen is devoid of ambulatory appendages, and the genital opening is situated near the posterior end of the body. Postembryonic development is rarely direct and a metamorphosis usually occurs."